+27 51 447 9609
36 Aliwal Street
Bloemfontein, South Africa

Terrestrial Invertebrates

Terrestrial Invertebrates

Study of Insects | Mites | Spiders

The Department Terrestrial Invertebrates consists of three collections: Acarology, Arachnology and Entomology. The research focus of the department involves projects in these three collections and the specialists working on it.

Currently the focus is:

  1. Taxonomy of South African oribatid mites
  2. Taxonomy of South African pseudoscorpions
  3. Taxonomy and systematics of Southern African scarab beetles
  4. Taxonomy and systematics of Afrotropical flies
Staff

Principal Museum Scientist and Head of Department - Acarology

Lizel Hugo-Coetzee PhD lhugo@nasmus.co.za

Lizel Hugo-Coetzee obtained her BSc, BSc Hons and MSc at the University of the Free State, majoring in Entomology. She obtained her PhD in Conservation Ecology at Stellenbosch University in 2006. Her thesis title was: Spatial patterns in the microarthropod community associated with Azorella selago (Apiaceae) on the sub-Antarctic Prince Edward Islands.

She started working at the National Museum in January 2006 as a Museum Scientist. Lizel is a Research Associate of the Department of Zoology and Entomology at the University of the Free State.

Lizel has been appointed Head of Department in January 2017. Her main interest is the taxonomy of oribatid mites in South Africa, as well as some ecological aspects of oribatid mite communities.


Principal Museum Scientist - Entomology

Gimo M. Daniel PhD (Pr.Nat.Sci.) gimo.daniel@nasmus.co.za

Gimo holds a BSc in Biology, MSc and PhD in Entomology. He obtained his PhD degree at the University of Pretoria in 2019, under supervision of Clarke Scholtz, Catherine Sole and Adrian Davis.  Gimo started working at the National Museum in July 2019 as Principal Museum Scientist. He also is a research fellow associated with the Department of Biological & Environmental Sciences at Walter Sisulu University.

Gimo is an entomologist with interests in understanding the systematics and biogeography of dung beetles and related scarab beetles. He is cataloguing and describing the impressive biodiversity of Afrotropical dung beetles to address broader evolutionary questions, such as the role of geological uplift and climatic changes in the late Cenozoic in the diversification and possible extinction of scarab beetles in southern Africa.


Senior Museum Scientist - Entomology

Burgert Muller M.Sc. burgert.muller@nasmus.co.za

B.Sc Entomology (UFS); M.Sc. Entomology(UFS)

Burgert MullerBurgert began his entomology career in 2008 as Senior Research Technician: Natural Sciences at the KwaZulu-Natal Museum, Pietermaritzburg. From 2009–2016 he was employed at the same institution as Curator of Entomology, focussing his research on Afrotropical Diptera, specifically Muscidae. During his time at the museum he served as Graphics Editor for the International (and museum) journal African Invertebrates. In 2014 he took over as Managing Editor of African Invertebrates until his departure from the Museum. He still acts as subject editor for the Taxonomy discipline in the journal. He also served as co-facilitator and trainer of Georeferencing workshops for SANBI since 2012.

In 2014 Burgert obtained his Masters degree for a thesis titled: Systematics of the shoot fly subgenus Atherigona s. str. (diptera: muscidae) of South Africa.

Burgert’s research interests are systematics, taxonomy, phylogeny and biogeography of true flies (Diptera), with special emphasis on Muscidae and Athericidae. He is also interested in Cybertaxonomy and literature mark-up, as well as Collections data quality assessment and use, which includes georeferencing and collections information management.


Museum Scientist - Arachnology

Jan Andries Neethling MSc ja.neethling@nasmus.co.za

Jan_AndriesJan registered for B.Sc Microbiology at the University of the Free State in 2006, obtaining the degree with distinction in 2009. In 2010 he registered for third year B.Sc Entomology, obtaining distinctions in all the registered subjects and was consequently accepted into the Honours program at the Zoology and Entomology department in 2011 where he specialised in Arachnid taxonomy, but also did courses in Forensic Entomology and Plant-Insect Interactions.

At the end of 2011 he obtained his Honours (cum laude) with his thesis: “Effect of moon phases on the activity patterns of trapdoor spiders, scorpions and solifugids at the Free State National Botanical Gardens”.

He recently (2015) obtained his Master of Science (cum laude) with his thesis: “Revision of the South African Geogarypidae (Arachnida: Pseudoscorpiones)”, which represents the first holistic taxonomic study of a South African pseudoscorpion family, taking both molecular and morphological characters into consideration.

He is currently busy with the taxonomic revision of the South African Pseudoscorpionida fauna.


Chief Research Assistant - Arachnology

Trudie-PeyperTrudie Peyper trudie@nasmus.co.za

Trudie Peyper is a National Museum Staff Member in the Arachnology Department

Trudie Peyper grew up in Bloemfontein. She completed a B Agric degree at the University of the Free State. She started to work at the Museum in 1984 and is currently a Chief Research Assistant in the Arachnology Department.


Mabocha Jarmaine Magoai – Research Assistant

Jarmaine obtained a B.Sc. degree in Entomology at the University of Pretoria and is currently an intern working in the Entomology collection. She is tasked with data basing of specimen data, verifying captured data, and general curation of specimens

Growing up, Jarmaine wanted to know about the dynamics of the environment, hence the field of Science. She loves nature and would like to further her studies in conservation.

 


Sarah Mokwena – Intern Data Capturer: Entomology

Sarah obtained her BSc in Entomology and Microbiology and BSc Hons in Entomology at the University of Fort Hare. She is tasked with data basing insect specimens in the Entomology collection.

From a young age she has always been curious how animals as small as insects could have such a vast impact on the environment and its ecosystem. She would like to continue her studies focusing mainly on Insect ecology.

Move to top
Current Research Projects

Project 556: A revision of the Hermannielliidae (Acari, Oribatida) of South Africa.

Project 555: Effects of climate and topography on spatial patterns of genetic structure in southern African dung beetles.

Project  457: Systematics of Afrotropical Athericidae (water snipe flies).

Project 542 Diversification and expansion of the National Museum Entomology collection

Project 504: Systematics of the endemic, Afrotropical, dung beetle genus,  Odontoloma (Scarabaeidae: Scarabaeinae)

Project 505: Revision of the South African pseudoscorpion fauna (Arachnida: Pseudoscorpiones).

Publications

Scientific publications

EA Hugo-Coetzee

  • Hugo-Coetzee, E.A. Notes on the family Oripodidae (Acari, Oribatida) in South Africa and description of a new species of Cryptoribatula Jacot. Systematic and Applied Acarology, 28(2), 394-404.
  • Ermilov, S.G., Hugo-Coetzee, E.A., Rybalov, L.B. 2023. Contribution to the knowledge of the oribatid mite genus Aleurodamaeus 9Acar, Oribatida, Aleurodamaeidae), with description of a new species from Ethiopia. Zoologicheskii Zhurnal, 102(2), 153-162.
  • Pfingstl, T., Wagner, M., Baumann, J., Neethling, J.A., Bardel-Kahr, I. & Hugo-Coetzee, E.A. 2022. Contrasting phylogeographic patterns of intertidal mites (Acari, Oribatida) along the South African shoreline. Organisms Diversity & Evolution, 22, 789-801.
  • Hugo-Coetzee, E.A., Baumann, J., Neethling, J.A., Bardel-Kahr, I. & Pfingstl, T. 2022. Ontogeny of South African intertidal oribatid mite species (Acari, Oribatida, Ameronothroidea) and supplements to adult morphology. Acarologia, 62(3), 721-753
  • Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov A.A. Khaustov V.A. 2021. A new species of Malgacheliodes (Acari, Oribatida, Licnodamaeidae) from South Africa. Zootaxa, 4984(1), 357-367. 
  • Ermilov, S.G., Hugo-Coetzee, E.A., Behan-Pelletier, V.M.  2021. Hogsbackia africaensis gen. nov., sp. nov. (Acari, Oribatida, Ceratozetoidea, Punctoribatidae), from an Afromontane forest of South Africa. Zootaxa, 4963 (3), 530-544.
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov, A.A. 2021. Contribution to the knowledge of Geminoppia (Acari, Oribatida, Oppiidae) with description of a new species from South Africa. Acta Zoologica Academiae Scientarum Hungaricae, 67(3), 211-222.
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. & Khaustov V.A. 2021. New faunistic data on oribatid mites (Acari, Oribatida) inhabiting the soil and termite nests in the Franklin Game Reserve, Bloemfontein, South Africa. Zoologicheskii Zhurnal, 100(4), 374-384.
  • Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov A.A. New species of oribatid mites of the subfamily Brachioppiinae (Acari, Oribatida, Oppiidae) from Hogsback, South Africa. Systematic and Applied Acarology, 26(4), 684-697.
  • Niedbala, W., Hugo-Coetzee, E.A. & Ermilov, S.G. 2021. New faunistical and taxonomic data on ptyctimous mites (Acarina, Oribatida) of South Africa. Systematic and Applied Acarology, 26(7), 1327-1349.
  • Pfingstl, T., Baumann, J., Neethling, J.A., Bardel-Kahr, I, Hugo-Coetzee, E.A. 2021. Distribution patterns of the oribatid intertidal mites (Acari, Oribatida) from South African shores and their relationship to temperature. African Journal of Marine Science, 43(2), 215-225.
  • Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov A.A. 2021. Three new species of oribatid mites of the family Galumnidae (Acari, Oribatida) from South Africa. Zootaxa 4920 (1), 091-100
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. & Khaustov, V.A. 2021. New faunistic and taxonomic data on oribatid mites (Acari, Oribatida) inhabiting soil and termite nests in the Golden Gate National Park (eastern South Africa). International Journal of Acarology. Doi.org/10.1080/01647954.2021.1879262
  • Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov A.A. 2021. A new species of Scheloribates (Acari, Oribatida, Scheloribatidae) from South Africa. Ecologica Montenegrina 39, 86-91.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2020. A new species of oribatid mites of the gemus Galumna from South Africa. Acarina 28(1), 17-22
  • Ermilov S.G. & Hugo-Coetzee E.A. 2020. Correspondence. Galumna capensis dissimilis Engelbrecht, 1969, a junior synonym of Galumna lawrencei Jacot, 1940 (Acari, Oribatida, Galumnidae), with remarks on type material. Zootaxa 4750(3): 441-442
  • Khaustov, A.A., Hugo-Coetzee, E.A., Ermilov, S.G. 2020. A new species of Lorryia (Acari: Tydeidae) from termite nest in South Africa. Acarina 28(1): 47-53
  • Joharchi, O., Hugo-Coetzee, E.A., Ermilov, S.G., Khaustov, A.A. 2020. Redescription of Hypoaspisella spiculifer (Berlese, 1918) comb. nov. (Acari: Mesostigmata: Laelapidae) from South Africa. Acarina 28(1); 55-64
  • Joharchi, O., Hugo-Coetzee, E.A., Ermilov, S.G., Khaustov, A.A. 2020. A new unusual species of Gaeolaelaps Evans and Till (Mesostigmata: Laelapidae) from South Africa. Annales Zoologici 70(3): 439-447
  • Khaustov, A.A., Hugo-Coetzee, E.A. & Ermilov S.G. 2020. Two new myrmecophilous Scutacarus (Acari: Scutacaridae) from South Africa. Annales Zoologici 70(3): 397-407
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. & Khaustov V.A. 2020. Ontogenetic instars of Phyllhermannia goldengatensis sp.nov. (Acari, Oribatida, Hermanniidae) from South Africa. Systematic and Applied Acarology 25(11): 2115-2130
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. 2020. Contribution to the knowledge of the oribatid mite genus Setoppia (Acari, Oribatida, Oppiidae), with a new species from South Africa. Acarologia 60 (4): 892-902
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. 2020. A new species of oribatid mites of the subgenus Lanceoppia (Bicristoppia) (Acari: Oribatida: Oppiidae) from South Africa. Zoosystematica Rossica 29(2), 328-334.
  • Khaustov, AA; Hugo-Coetzee, EA; Ermilov SG (2019) A new species of Tanytydeus (Acari: Paratydeidae) from termite nests in South Africa. Systematic and Applied Acarology 24(9), 1604-1619
  • Ermilov, SG; Hugo-Coetzee, EA; Khaustov, AA; Theron, PD (2019) Oribatid mites (Acari, Oribatida) inhabiting termite nests in the Faan Meintjes Nature reserve (South Africa). Systematic and Applied Acarology 24(9), 1783-1798
  • Khaustov, AA; Hugo-Coetzee, EA; Ermilov, SG (2019) A new genus, new species and a new record of the family Pygmephoridae (Acari: Heterostigmata) associated with Microtermes parvus (Havialnd) (Isoptera: Termitidae) from South Africa. Systematic and Applied Acarology 24(10): 1881-1892
  • Ermilov, SG; Hugo-Coetzee, EA; Khaustov, AA; Kontschan, J (2019) Hypozetes andreii (Acari, Oribatida, Tegoribatidae), a new species of oribatid mites from South Africa. Acarina 27(2): 183-192
  • Niedbała, W., Hugo-Coetzee, E.A. & Ermilov, S.G. 2019. New Notophthiracarus species (Acarina, Oribatida, Phthiracaridae) and overview of the distribution of the genus in South Africa. Zootaxa 4647 (1): 231-240
  • Khaustov, A.A., Hugo-Coetzee, E.A., Ermilov, S.G. & Theron, P.D. 2019. A new genus and species of the family Microdispidae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Sjöstedt) (Isopteran: Termitidae) from South Africa. Zootaxa 4647 (1): 104-114
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2019. New data on oribatid mites (Acari, Oribatida) of South Africa, with description of two new species of the family Oppiidae. Systematic and Applied Acarology 24(2): 287-302
  • Bayartogtokh, B., Ermilov, S.G., Hugo-Coetzee, E.A. Khaustov, A.A. 2018. Contribution to the knowledge of the oribatid mites genus Licnodamaeus Grandjean, 1931 and synonymy of Licnodamaeolus Covarrubias, 1998 (Acari, Oribatida, Licnodamaeidae). Systematic and Applied Acarology 23(1): 42-60
  • Ermilov, S.G., Hugo-Coetzee, E.A.; Theron, P.D. 2018. To the knowledge of oribatid mites of the subgenus Galumna (Galumna) Heyden 1826 (Acari, Oribatida, Galumnidae) in South Africa, with a key to species known from the Ethiopian region. Zoologichesky Zhurnal 97(5): 515-527.
  • Khaustov, A.A., Hugo-Coetzee, E.A. & Ermilov, S.G. 2018. A new species of Scutacarus (Acari: Heterostigmata: Scutacaridae) associated with Trinervitermes trinervoides (Isoptera: Termitidae) from South Africa. International Journal of Acarology, online
  • Ahaniazad, M., Bagheri, M., Akrami, M.A. & Hugo-Coetzee, E.A. 2017.  Ameroid mites (Acari; Oribatida) from northwest of Iran with description of a new species of Ctenobelba Balogh, 1943. Systematic & Applied Acarology 22(1): 74–84.
  •  Ermilov, S.G., Hugo-Coetzee, E.A. Khaustov, A.A. 2017.  Oribatid mites (Acari, Oribatida) inhabiting nests of the termite Trinervitermes trinervoides (Sjöstedt) in the Franklin Game Reserve (Bloemfontein, South Africa), with description of a new species of the genus Ceratobates (Tegoribatidae). Systematic and Applied Acarology 22(10): 1715-1732
  • Ermilov, S.G., Hugo-Coetzee, E.A., Theron, P.D. 2017. New and interesting moss mites (Acari, Oribatida) from South Africa, with description of two new species. Systematic and Applied Acarology 22(10): 1560-1573
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2017. New data on oribatid mites of the family Suctobelbidae (Acari: Oribatida) from South Africa, with description of a species of the genus Suctobelbella. Biologia 72(5): 536-541
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. & Theron, P.D. 2017. New and interesting oribatid mites (Acari, Oribatida) near Potchefstroom (South Africa), with description of two new species. Systematic and Applied Acarology 22(11): 1849–1871.
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov A.A. 2017. Contribution to the knowledge of the oribatid mite genus Lamellarea (Acari, Oribatida, Lamellareidae). Systematic & Applied Acarology 22(11): 2008–2022.
  • Ermilov, S.G., Hugo-Coetzee, E.A.; Theron, P.D.; Behan-Pelletier, V.M. 2017. Contribution to the knowledge of the oribatid mite family Nesozetidae (Acari, Oribatida). Zootaxa 4358(2): 311-327.
  • Ermilov, S.G., Hugo-Coetzee, E.A., Khaustov, A.A.; Theron, P.D. 2017. Juvenile instars of Galumna curvifamulus  (Acari, Oribatida, Galumnidae). Acarina 25 (2): 87-95
  • Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.  Oribatid mites (Acari, Oribatida) inhabiting nests of the termite Trinervitermes trinervoides (Sjöstedt) in the Franklin Game Reserve (Bloemfontein, South Africa), with description of a new species of the genus Ceratobates (Tegoribatidae). Systematic and Applied Acarology 22(10): 1715-1732.
  • Ermilov, S.G.; Hugo-Coetzee, E.A.; Theron, P.D.2017. New and interesting moss mites (Acari, Oribatida) from South Africa, with description of two new species. Systematic and Applied Acarology 22(10): 1560-1573.
  • Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Contribution to the knowledge of oribatid mites of the family Lohmanniidae (Acari: Oribatida) from South Africa. Systematic and Applied Acarology 22(5): 666-682.
  • Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Two new species of the genus Ausoribula (Acari, Oribatida, Oribatulidae) from termitaries of South Africa. Acarologia 57(3): 643-650.
  • Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017. Coetzeella navalensis gen. nov. sp. nov. (Acari; Oribatida; Oppiidae) from South Africa.Systematic & Applied Acarology 22(3): 403-409.  
  • Hugo-Coetzee, E.A.2017. New Oppiidae (Acari: Oribatida) from Vernon Crookes Nature Reserve, South Africa. Zootaxa 4311(2): 211-232.
  • Hugo-Coetzee, E.A.; Lotfollahi, P.; Abbasi-Kalo, A.; Movahedzade, E.2017. A new species of Oxyoppia (Oxyoppiella) (Acari, Oribatida, Oppiidae) from Iran. Turkish Journal of Zoology 41: 630-637.
  • Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017.A new genus and two new species of Pygmephoridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: termitidae) from South Africa. Systematic and Applied Acarology 22(4): 484-493.
  • Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017. A new genus of the mite family Scutacaridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: Termitidae) from South Africa. Zootaxa 4258(5): 462-476.
  • Hugo-Coetzee, E.A. 2016.  New species of Ramusella (Oribatida, Oppiidae) from South Africa with additional notes on R. filamentosa (Mahunka) and R. filigera(Mahunka) Systematic & Applied Acarology,21(2): 227–244
  • Janion-Scheepers, C. et al. 2016. Soil biota in a megadiverse country: Current knowledge and future research directions in South Africa. Pedobiologia 59: 129-174.
  • Hugo-Coetzee, E.A. 2014. A new species of Adrodamaeus (Acari, Oribatida, Gymnodamaeidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 30(6): 87-99.
  • Hugo-Coetzee, E.A. 2014. New Oppiidae (Acari: Oribatida) from the Golden Gate Highlands National Park in South Africa. Zootaxa 3884(6): 533-552.
  • Hugo-Coetzee, E.A. 2013. New species of Aleurodamaeus Grandjean, 1954 (Oribatida: Aleurodamaeidae) from South Africa. Zootaxa 3670(4): 531-556.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species of Nothrus (Acari: Oribatida: Nothridae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 28(2): 25-40.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Valbehanella freestatensis gen. nov., sp. nov. (Acari, Oribatida) from South Africa.  Systematic & Applied Acarology 17(4): 407-416.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Oribatid mites of the genus Eremulus Berlese, 1908 (Acari: Oribatida: Eremulidae) from South Africa.  African Invertebrates 53(2): 559-569.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. The oribatid mite genus Nothrus Koch, 1836 (Acari: Oribatida: Nothridae) of South Africa, including a key to African species.  Zootaxa 3243: 29-51.
  • Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species from South Africa, with remarks on generic diagnosis of Licnodamaeolus Covarrubias, 1998 and taxonomic status of Nacunansella Fernandez & Cleva, 1998 (Acari: Oribatida: Licnodamaeidae). Zootaxa 3167: 32-44.
  • Hugo-Coetzee, E.A. & Avenant, N.L. 2011. The effect of fire on soil oribatid mites (Acari: Oribatida) in a South African grassland.  Zoosymposia 6: 210-220.
  • Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov, A.A. 2011. Morphology of juvenile instars of Neoliodes terrestris (Wallwork, 1963) and N. ionicus Sellnick, 1931 (Acari: Oribatida: Neoliodidae).  Annales Zoologici (Warszawa) 61(4): 817-830.
  • Hugo-Coetzee, E.A. 2011.  Three new species of Austrocarabodes (Oribatida: Carabodidae) and notes on Austrocarabodes pinnatus Mahunka, 1986, from South Africa.  Zootaxa 3011: 1-15. 
  • Hugo-Coetzee, E.A. 2010. Two new species of Austrocarabodes (Acari: Oribatida: Carabodidae) from South Africa: A. mahunkai and A. lineasetosaNavorsinge van die Nasionale Museum, Bloemfontein 26(2): 45-58.
  • Hugo, E.A. 2010. Two new species of Gymnodamaeidae (Acari: Oribatida) from South Africa.  International Journal of Acarology  36(3): 199-210. 
  • Hugo, E.A. 2008. Three new species of Austrocarabodes (Oribatida: Carabodidae) from South Africa. Zootaxa 1844: 25-36. 
  • Hugo, E.A., Chown, S.L. & McGeoch, M.A. 2006. The microarthropods of sub-Antarctic Prince Edward Island: a quantitative assessment. Polar Biology, 30(1):109-119 
  • Hugo, E.A., McGeoch, M.A., Marshall, D.J. & Chown, S.L. 2004. Fine scale variation in microarthropod communities inhabiting the keystone species Azorella selago on Marion Island. Polar Biology, 27(8):466-473. 


Gimo M. Daniel

  • Nganhane, I.de N., Naskrecki, P., Farooq, H., Daniel, G.M. Millipede consumption and interaction between food preferences and sex in dung beetle Chalconotus convexus (Coleptera:Scarabaeinae) Austral Ecology
  • Daniel, G. M., & Davis, A. L. 2023. Dung beetle tribal classification (Coleoptera: Scarabaeidae: Scarabaeinae): progress, problems, and prospects. Annals of the Entomological Society of America, saad032.
  • Daniel, G. M., Josso, J. F., Nganhane, I. & Strumpher, W. P. 2024. Dung beetle fauna from Mount Mabu, Mozambique. Part 3: A new species of Proagoderus van Lansberge, 1833 with an annotated checklist of species of Proagoderus dives (Harold, 1877) species-group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 5406, 175–189.
  • Stals, R., Daniel, G. M. & Deschodt, C. 2024. Hathoronthophagus, new replacement name for Hathor Deschodt, 2023, preoccupied genus-group name of a putatively myrmecophilic dung beetle (Scarabaeidae: Scarabaeinae: Onthophagini). Zootaxa, 5397, 449–450
  • Daniel, G.M., Strümpher, W.P., Josso, J.-F. 2023. Dung beetle fauna from Mount Mabu, Mozambique. Part 1: A new species of Onthophagus Latreille, 1802, and a checklist of species belonging to d'Orbigny's group 3 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 5258 (4): 429–442.

  • Daniel, G.M., Strümpher, W.P., Josso, J.-F. 2023. Dung beetle fauna of Mabu. Part 2: Pedaria ricardogutai (Coleoptera: Scarabaeidae), a new species, with an annotated checklist of species of Pedaria Laporte, 1832 from Mozambique. Zootaxa 5325(3), 409-418.

  • Biffi, G., Leschen, R. A. B., Hsiao, Y., Daniel, G. M., Casari, S. A. 2022. The systematics of Dysmorphocerinae (Cantharidae) based on larvae. Insect Systematics & Evolution (published online). doi: https://doi.org/10.1163/1876312X-bja10041
  • Dubois, A., Aescht, E., Aneesh, P. T., Ceríaco, L. M. P., Daniel, G. M., De Prins, J., Engel, M.S., Frétey, T., Löbl, I., Lorvelec, O., Marinov, M., Ohler, A., Schmitt, M., Whittington, A., Young, M.T., Bauer, A. M. 2022. The Linz Zoocode project. Sixth report of activities. Nomenclatural availability. 5. Optical disc publication. Bionomina31(1), 24-45.
  • Strumpher, W., Daniel, G.M., Davis, A.L.V. 2022. Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolow & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 5188(4), 396-100.
  • Daniel, G. M., Strümpher, W. P. & Deschodt, C. M. 2022. An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South AfricaZootaxa, 5093(4), 414–420.
  • Daniel, G.M., Noriega, J.A., da Silva, P.G., Deschodt, C.M., Sole, C.L., Scholtz, C.H. & Davis, A.L.V. 2022. Soil type, vegetation cover and temperature determinants of the diversity and structure of dung beetle assemblages in a South African open woodland and closed canopy mosaic. Austral Ecology, 47: 79-91.
  • Dubois, A., Aneesh, P.T., Bauer, A.M., Ceríaco, L.M.P., Daniel, G.M., Frétey, T., Löbl, I., Lorvelec, O., Marinov, M., Ohler, A., Schmitt, M., Whittington, A. & Aescht, E. 2022. The Linz Zoocode project. Second report of activities (2020). Nomenclatural availability. 1. What is nomenclatural availability? Bionomina, 28, 1–17.
  • Dubois, A., Aneesh, P.T., Bauer, A.M., Ceríaco, L.M.P., Daniel, G.M., Frétey, T., Löbl, I., Lorvelec, O., Marinov, M., Ohler, A., Schmitt, M., Whittington, A. & Aescht, E. 2022. The Linz Zoocode project. Third report of activities (2020). Nomenclatural availability. 2. Work availability. Bionomina, 28, 18–49.
  • Dubois, A., Aneesh, P.T., Bauer, A.M., Ceríaco, L.M.P., Daniel, G.M., De Prins, J., Frétey, T., Löbl, I., Lorvelec, O., Marinov, M., Ohler, A., Schmitt, M., Whittington, A., Young, M. T. & Aescht, E. 2022. The Linz Zoocode project. Fourth report of activities (2021). Nomenclatural availability. 3. Paper publication. Bionomina, 28, 50–70.
  • Dubois, A., Aneesh, P.T., Bauer, A.M., Ceríaco, L.M.P., Daniel, G.M., De Prins, J., Frétey, T., Löbl, I., Lorvelec, O., Marinov, M., Ohler, A., Schmitt, M., Whittington, A. & Aescht, E. 2022. The Linz Zoocode project. Fifth report of activities (2022). Nomenclatural availability. 4. Electronic publication. Bionomina, 28, 71‒119.
  • Strűmpher, W., Daniel, G.M., Davis, A.L.V. (2022) Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolow & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 5188(4), 396-100.
  • Daniel, G.M., Sole, C.L., Scholtz, C.H., Davis, A.L.V. 2021. Historical diversification and biogeography of the endemic southern African dung beetle genus, Epirinus (Scarabaeidae: Scarabaeinae). Biological Journal of the Linnean Society, blab051.b 
  • Strümpher, W.P., Daniel, G.M. 2021. A new dung beetle species of the genus Stiptopodius Harold, 1871 (Coleoptera: Scarabaeidae: Scarabaeinae) recorded from xeric savanna in Namibia. Zootaxa, 4999(4), 335-342.
  • Engel, M.S., Ceríaco, M.L.P., Daniel, G.M et al. 2021. The taxonomic impediment: a shortage of taxonomists, not the lack of technical approaches. Zoological Journal of the Linnean Society, 193(2) 381–387.
  • Daniel, G. M., Strümpher, W. P., Snäll, S. 2021. A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa. Zootaxa5072(1), 34-42. Dubois, A., Panakkool, T.A., Aaron, M.B, Ceríaco, .L.P, Daniel, G.M., Dellapé, P., Engel M.S. et al. 2021.  Optical discs in zoological nomenclature: problems and proposed solution. Bionomina 24, (1) 1-8.
  • Daniel, GM., Davis, ALV, Sole, CL, & Scholtz, CH. 2020. Evolutionary history and eco-climatic diversification in southern African Sisyphus (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Biogeography; 47: 2698-2713. https://doi.org/10.1111/jbi.13974
  • Daniel, GM., Deschodt, CM., Davis, ALV., & Sole, CL. 2020. The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera: Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa. Zootaxa, 4763(3), 394-404.
  • Daniel, GM, Sole, CL, Davis, ALV, Strumpher, WP & Scholtz, CH. 2020.  The dung beetle tribe Sisyphini Mulsant sensu novo (Scarabaeinae: Scarabaeidae): assessment of evolution and systematics inferred from molecular phylogeny and geographical distribution of southern African species. Systematic Entomology, 73-84.
  • Daniel, GM., Davis, ALV, Sole, CL, & Scholtz, CH. 2020. Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions, Insect Systematic and Evolution, 1-61.
  • Daniel, GM The nomenclatural status of the generic nomina Epirinus Dejean, 1833 and Epirhinus Agassiz, 1846 (Coleoptera, Scarabaeidae, Scarabaeinae), Bionomina, 63-65.
  • Daniel, GM & Génier, F 2019. Checklist of the dung beetle species of Mozambique (Coleoptera: Scarabaeidae: Scarabaeinae), Catharsius, La Revue, 25-32.
  • Deschodt, CM., Davis, ALV., & Daniel, GM. (2019). The endemic Southern African genus Epirinus Dejean, 1833 (Coleoptera: Scarabaeidae: Scarabaeinae) revisited: five new species and two new synonyms. Zootaxa, 4603(2), 327-340.
  • Daniel, GM, Cupello, M. Ateuchus hirtusWiedmann 1823, (Insecta, Coleoptera, Scarabaeidae) proposed conservation of the specific name, Bulletin of Zoological Nomenclature, 123-126.
  • Daniel, GM., Davis, ALV., & Scholtz, CH. 2016. Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa.Zootaxa4147, 67-74.
  • Daniel, GM & Vaz-de-Mello, FZ. 2016. Biotic components of dung beetles (Insecta: Coleoptera: Scarabaeidae: Scarabaeinae) from Pantanal–Cerrado Border and its implications for Chaco regionalization. Journal of Natural History 50, 1159–1173.
  • Daniel GM; Nunes LGOA & Vaz-de-Mello FZ. 2014. Species composition and functional guilds of dung beetles (Insecta: Coleoptera: Scarabaeidae: Scarabaeinae) in different vegetational types in the Brazilian Shield-Chacoan Depression Border. Annales de la Societe entomologique de France. 50, 183-190.


Burgert S. Muller

  • Midgley, J.M., Muller, B.S. Description of the female of Atherimorpha latipennis Stuckenberg (Diptera, Rhagionidae): the first record of brachyptery in Rhagionidae. ZooKeys, 1178, 265-277.
  • Midgley, J.M., Muller, B.S., Theron, G.L., Phoofolo M., Bellingan, T.A., Jordaens, K. (2023) The Diptera of Lesotho: a history of collecting in the Mountain Kingdom, summary of recent collecting sites and introduction to the tropical collection in African Invertebrates. African Invertebrates, 64(3), 207-220.
  • Muller, B.S., Swart, V.R., Snyman, L.P. 2023. Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species. African Invertebrates, 64(3), 303-322.
  • Muller, B.S., Midgley, J.M. 2022. How strange: Coenosia curiosa nov. (Diptera: Muscidae), the first recorded Tiger fly from Lesotho, with revision of the Coenosia globuliseta-group. Zootaxa 5222(4): 367-377.
  • Demetriou, J., Kalaentzis, K., Kazilas, C., Kunz, G., Muller, B., Mostovski, M.B. & Koutsoukos, E. 2022. An “alien” species on the loose: new records and updated distribution of the black soldier fly Hermetia illucens in the Western Palearctic. Bulletin of Insectology, 75(1), 125-130.
  • Muller, B.S. 2021. Piophilidae (Skipper Flies) 68. In: Kirk-Spriggs, A.H. & Sinclair, B.J. (eds) Manual of Afrotropical Diptera. Volume 3. Brachycera-Cyclorrhapha, excluding Calyptratae. Suricata 8. South African Biodiversity Institute, Pretoria: pp. 1597-1606
  • Muller, B.S. 2021. Braulidae (Bee Louse Flies) 105. In: Kirk-Spriggs, A.H. & Sinclair, B.J. (eds) Manual of Afrotropical Diptera. Volume 3. Brachycera-Cyclorrhapha, excluding Calyptratae. Suricata 8. South African Biodiversity Institute, Pretoria: pp. 2289-2294
  • Muller, B.S. 2019. A curious new Coenosia Meigen, 1826 (Diptera, Muscidae) from Mariepskop, Mpumalanga, South Africa. African Invertebrates 60(2): 239–253.
  • Muller, B.S. & Mostovski, M.B. 2018. A key to males of Atherigona s. str. (Diptera: Muscidae) from Mali, with new records and a new species for the country. Zootaxa 4425 (2): 342–356.
  • Kirk-Spriggs, A.H. & Muller, B.S. 2017. 9. Biogeography of Diptera. In: Kirk-Spriggs, A.H. & Sinclair, B.J., eds, Manual of Afrotropical Diptera. Volume 1. Introductory chapters and keys to Diptera families. Suricata 4. Pretoria: SANBI Graphics & Editing, 203–238.
  • Marshall, S.A., Kirk-Spriggs, A.H., Muller, B.S., Paiero, S.M., Yau, T. & Jackson, M.D. 2017. 12. Key to Diptera families—adults. In: Kirk-Spriggs, A.H. & Sinclair, B.J., eds, Manual of Afrotropical Diptera. Volume 1. Introductory chapters and keys to Diptera families. Suricata 4. Pretoria: SANBI Graphics & Editing, 267–355. 
  • Muller, B.S. 2015. Illustrated Key and Systematics of Male South African Atherigona s. str. (Diptera: Muscidae). African Invertebrates 56 (3): 845–918.
  • Muller, B.S. 2013. Rediscovering the Old from New: Two Curious Species of Coenosia Meigen (Diptera: Muscidae) from South Africa. African Invertebrates 54 (2): 595–603.
  • Miller, J., Dikow, T., Agosti, D., Sautter, G., Catapano, T., Penev, L., Zhang, Z.Q., Pentcheff, D., Pyle, R., Blum, S., Parr, C., Freeland, C., Garnett, T., Ford, L.S., Muller, B., Smith, L., Strader, G., Georgiev, T., Bénichou, L. From taxonomic literature to cybertaxonomic content. 2012. BMC Biology 10 (87): 1–5.
  • Oliveira, S.S. & Muller, B.S. 2012. The types of Lygistorrhinidae and Mycetophilidae (Diptera: Bibionomorpha) in the KwaZulu-Natal Museum, Pietermaritzburg, South Africa. African Invertebrates 53 (2): 703–714.
  • Davies, G.B.P., Miller, R.M. & Muller, B.S. 2012. A new genus of lauxaniid fly from South Africa (Diptera: Acalyptratae: Lauxaniidae), associated with proteas (Proteaceae). African Invertebrates 53 (2): 615–636.
  • Grichanov, I.Y., Mostovski, M.B. & Muller, B.S. 2011. New records of Afrotropical Dolichopodidae (Diptera) from the Collection of the Natal Museum (2). International Journal of Dipterological Research 22 (2): 65–82.
  • Grichanov, I.Y., Mostovski, M.B. & Muller, B.S. 2011. New records of Afrotropical Dolichopodidae (Diptera) from the Collection of the Natal Museum (1). International Journal of Dipterological Research 22 (1): 3–9.


Jan A. Neethling

  • Haddad, C.R., Prendini, L., Neethling, J.A. & Dippenaar-Schoeman, A.S. 2023. The non-acarine Arachnida of the Amathole Mountains, South Africa. Bothalia, 53(1), 1-33.
  • Neethling, J.A. & Neethling, C. 2023. A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea). Zootaxa, 5256(6), 501-543.

  • Pfingstl, T., Wagner, M., Baumann, J., Neethling, J.A., Bardel-Kahr, I. & Hugo-Coetzee, E.A. 2022. Contrasting phylogeographic patterns of intertidal mites (Acari, Oribatida) along the South African shoreline. Organisms Diversity & Evolution, 27, 789-801.
  • Hugo-Coetzee, E.A., Baumann, J., Neethling, J.A., Bardel-Kahr, I. & Pfingstl, T. 2022. Ontogeny of South African intertidal oribatid mite species (Acari, Oribatida, Ameronothroidea) and supplements to adult morphology. Acarologia 62(3), 721-753.
  • Neethling, J.A. & Haddad, C.R 2019. Influence of some abiotic factors on the activity patterns of trapdoor spiders, scorpions and camel spiders in a central South African grassland. Transactions of the Royal Society of South Africa
  • Neethling, J.A. & Haddad, C.R. 2016. A systematic revision of the South African pseudoscorpions of the family Geogarypidae. Indago 32: 1-80.
  • Janion-Scheepers, C., Measeya, J., Braschlera, B., Chown, S.L., Coetzee, L., Colville, J.F., Dames, J., Davies, A.B., Davies, S.J., Davis, A.L.V., Dippenaar-Schoeman, A.S., Duffy, G.A., Fourie, D., Griffiths, C., Haddad, C.R., Hamer, M., Herbert, D.G., Hugo-Coetzee, E.A., Jacobs, A., Jacobs, K., Jansen van Rensburg, C., Lamani, S., Lotz, L.N., Louw S. vdm., Lyle, R., Malan, A.P., Marais, M., Neethling, J.A., Nxele, T.C., Plisko, D.J., Prendini, L., Rink, A.N., Swart, A., Theron, P., Truter, M., Ueckermann, E., Uys, V.M., Villet, M.H., Willows-Munro, S., Wilson, J.R.U. 2016. Soil biota in a megadiverse country: Current knowledge and future research directions in South Africa. Pedobiologia 59: 129–174.
  • Neethling, J.A. & Haddad, C.R. 2013. Arboreal spider assemblages associated with four tree species in the Grassland Biome of central South Africa (Arachnida: Araneae). Transactions of the Royal Society of South Africa 68(2): 123-131.
  • Haddad, C.R., Neethling, J.A. & Lyle, R. 2011. Spinotrachelas montanus sp. n., the first Afromontane representative in the genus (Araneae: Corinnidae). African Invertebrates 52(2): 345-352.


Precious Tshililo

  • Tshililo, P. 2020. Review of South African Euryphyminae. Zootaxa 4820 (1), 070-104
  • Schoeman, C.S., Toussaint, D.C., Tshililo, P., Foord, S.H. and Hamer, M., 2020. Checklist of the pimeliine darkling beetles of the Vhembe Biosphere Reserve, South Africa (Coleoptera: Tenebrionidae: Pimeliinae). Bothalia-African Biodiversity & Conservation50(1), pp.1-9.
© 2019 National Museum, Bloemfontein, South Africa – All Rights Reserved
Website development by Digital Platforms