Acarologists appointed Research Associates

11 January 2017
Drs Louise Coetzee (left) and Lizel Hugo-Coetzee have been appointed Research Associates in the Faculty of Natural and Agricultural Sciences at the Univeristy of the Free State.  They will be collaborating with the Department of Zoology and Entomology.

EURAAC in Valencia, Spain

10 January 2017
Dr Lizel Hugo-Coetzee attended the 8th symposium of the European Association of Acarologists (EURAAC) in Valencia, Spain in July. There were 140 delegates from 33 countries. Topics included mite taxonomy, ecology, biological control, climate change, invasions and forensics. The title of Lizel’s presentation was Distribution patterns of microarthropods on sub-Antarctic Marion Island across altitude and aspect: climate change implications.

From left: Drs Lizel Hugo-Coetzee (South Africa), Andrea Lienhard (Austria), Julia Jagersbacher-Baumann (Austria), Satoshi Shimano (Japan) and Tobias Pfingstl (Austria).

Mites in Art

12 December 2014
Oliewenhuis Art Museum Education Officer, Yolanda de Kock, came up with the brilliant idea to “pair” departments at the National Museum with contemporary South African artists to observe the investigations, results and activities of researchers and use this information as inspiration to create artworks.

Oribatid mites found their way to two artworks – one by Ben Botma, “Tussen-in” (scraperboard and digital print), and one by Elrie Joubert, who made digital prints from light microscope images. For this project, soil was collected from the garden of Oliewenhuis Art Museum, and the oribatids were extracted, identified and photographed.

The project of combining research, education and exhibition fits the mission of the National Museum perfectly: to provide heritage resources and an enjoyable experience to all people through quality research, conservation, education and exhibitions.

The Inquisitive Mind: Science and Imagination can be viewed at Oliewenhuis Art Museum until 1 February 2015.

Second workshop of the Soil Ecosystem Research Group

19 September 2013
Louise Coetzee and Lizel Hugo-Coetzee of the Department of Acarology attended the second meeting of the Soil Ecosystem Research Group (SERG), established in 2011, aiming to form a network to improve communication and develop links between researchers involved in studies of soil biota. The second workshop of SERG was held during the 18th Congress of the Entomological Society of Southern Africa (ESSA XVIII) from 1 to 3 July at Potchefstroom. Progress on a paper to review the status of knowlegde regarding soil biota in South Africa was discussed. The paper is collaboratively drafted with researchers from various institutions and universities, all working on taxonomy of soil biota. Integrative sampling techniques and funding opportunities were also discussed.

EURAAC 2012

21 August 2012
Louise Coetzee attended the 7th symposium of the European Association of Acarologists (EURAAC), held from 9 – 13 July 2012 at the University of Natural Resources and Life Sciences (BOKU), Vienna, Austria. Louise presented a paper “An overview of the genus Afroleius (Acari, Oribatida)” in which certain character states not previously noticed in this genus were illustrated and a reinterpretation of other characters were discussed.

Apart from the scientific content of a conference, it is also a valuable experience to make social contact with colleagues from all over the world.

Louise Coetzee and Prof. Reinhardt Schuster, retired acarologist from Graz, Austria.

Soil Ecology Workshop

21 July 2011
A soil ecology workshop was held on 6 July 2011 during the 17th Congress of the Entomological Society of Southern Africa, held at the University of the Free State from 3 – 6 July. The workshop was attended by taxonomists and other researchers concerned with soil biota. Presentations were made by a number of specialists working on various ecologically important soil dwellers. The message from this workshop was clear: the shortage of taxonomists in these groups is severe. A soil ecology research group was established in order to find measures to address this problem.

Louise Coetzee presented a talk “Mites in Soil” pointing out the ecological role of oribatid (soil) mites as well as the taxonomic status of Oribatida in South Africa. Dr Lizel Hugo-Coetzee presented a poster (co-authored by Dr Nico Avenant) “Soil mite (Acari: Oribatida) response to prescribed fire in a central South African grassland”.

Attendants at the inaugural meeting of the Soil Ecology Research Group (Louise Coetzee 2^nd row, second from left; Lizel Hugo-Coetzee 2^nd row, third from left).

International Congress of Acarology

6 October 2010

Dr Lizel Hugo-Coetzee from the Acarology Department, National Museum, Bloemfontein, attended the 13th International Congress of Acarology from 23-27 August, 2010, in Recife, Brazil, where she reported on the effect of fire on soil mite communities in Free State grasslands.

The Congress was attended by 380 delegates from 41 countries. Other South Africans present at the Congress were Dr Eddie Ueckermann (Plant Protection Research Institute, Agricultural Research Council), Prof. Pieter Theron (School of Environmental Sciences & Development, North West University), Dr Louwrens Tiedt (Laboratory of Electron Microscopy, North West University) and Dr Jacob den Heyer (Department of Zoology & Entomology, University of the Free State.

Dr Lizel Hugo-Coetzee (4th from left) with participants at the Acarology Congress, Recife, Brazil

Principal Museum Scientist and Head of Department

Lizel Hugo-Coetzee PhD lhugo@nasmus.co.za

Lizel Hugo-Coetzee obtained her BSc, BSc Hons and MSc at the University of the Free State, majoring in Entomology. She obtained her PhD in Conservation Ecology at Stellenbosch University in 2006. Her thesis title was: Spatial patterns in the microarthropod community associated with Azorella selago (Apiaceae) on the sub-Antarctic Prince Edward Islands. Lizel spent a year on Marion Island collecting data for this degree. She started working at the National Museum in January 2006 as a Museum Scientist. Her main interest is the taxonomy of oribatid mites in South Africa, as well as some ecological aspects of oribatid mite communities. Lizel is a Research Associate of the Department of Zoology and Entomology at the University of the Free State. She is also a section editor for the journal, Systematic and Applied Acarology. Lizel has been appointed Head of Department in January 2017.

The objective of this Department is the taxonomic research of Oribatida (Acari) in order to apply our knowledge to control and sustain the condition of soil and the environment. Our main function is to describe the unknown oribatids of South Africa.

Current projects

Lizel Hugo-Coetzee
New species of the Family Oppiidae in South Africa.

Please contact the Acarology department for papers not available from this website.

E.A. Hugo-Coetzee lhugo@nasmus.co.za

Hugo-Coetzee, E.A.2017. New Oppiidae (Acari: Oribatida) from Vernon Crookes Nature Reserve, South Africa. Zootaxa 4311(2): 211-232.

Hugo-Coetzee, E.A.; Lotfollahi, P.; Abbasi-Kalo, A.; Movahedzade, E.2017. A new species of Oxyoppia (Oxyoppiella) (Acari, Oribatida, Oppiidae) from Iran. Turkish Journal of Zoology 41: 630-637.

Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.  Oribatid mites (Acari, Oribatida) inhabiting nests of the termite Trinervitermes trinervoides (Sjöstedt) in the Franklin Game Reserve (Bloemfontein, South Africa), with description of a new species of the genus Ceratobates (Tegoribatidae). Systematic and Applied Acarology 22(10): 1715-1732.

Ermilov, S.G.; Hugo-Coetzee, E.A.;Theron, P.D.2017. New and interesting moss mites (Acari, Oribatida) from South Africa, with description of two new species. Systematic and Applied Acarology22(10): 1560-1573.

Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017. A new genus of the mite family Scutacaridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: Termitidae) from South Africa. Zootaxa 4258(5): 462-476.

Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Contribution to the knowledge of oribatid mites of the family Lohmanniidae (Acari: Oribatida) from South Africa. Systematic and Applied Acarology 22(5): 666-682.

Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Two new species of the genus Ausoribula(Acari, Oribatida, Oribatulidae) from termitaries of South Africa. Acarologia 57(3): 643-650.

Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017.A new genus and two new species of Pygmephoridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: termitidae) from South Africa. Systematic and Applied Acarology 22(4): 484-493.

Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017. Coetzeella navalensis gen. nov. sp. nov. (Acari; Oribatida; Oppiidae) from South Africa.Systematic & Applied Acarology 22(3): 403-409.

Ahaniazad, M., Bagheri, M., Akrami, M.A. & Hugo-Coetzee, E.A. 2017.  Ameroid mites (Acari; Oribatida) from northwest of Iran with description of a new species of Ctenobelba Balogh, 1943. Systematic & Applied Acarology 22(1): 74–84.

Hugo-Coetzee, E.A. 2016.  New species of Ramusella (Oribatida, Oppiidae) from South Africa with additional notes on R. filamentosa (Mahunka) and R. filigera(Mahunka) Systematic & Applied Acarology,21(2): 227–244

Janion-Scheepers, C. et al. 2016. Soil biota in a megadiverse country: Current knowledge and future research directions in South Africa. Pedobiologia 59: 129-174.

Coetzee, L. 2015. Key to the species of Afroleius Mahunka, 1984 (Acari, Oribatida, Punctoribatidae), recombination of A. polygonatus (Mahunka, 1985), description of A. lucidus sp. nov. and discussion of A. undulatus (Balogh, 1959). Systematic & Applied Acarology 20(4): 383–398. view link

Hugo-Coetzee, E.A. 2014. A new species of Adrodamaeus (Acari, Oribatida, Gymnodamaeidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 30(6): 87-99. PDF 1.5 MEG

Coetzee, L. 2014. Rare new species of the genus Afroleius Mahunka, 1984 (Acari, Oribatida, Mycobatidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 30(5): 71-85. PDF 1.8 MEG

Coetzee, L. 2014. Afroleius floridus (Mahunka, 1985) comb. nov. and three new Afroleius Mahunka, 1984 species (Acari: Oribatida: Mycobatidae) from South Africa. Zootaxa (4): 553-573.

Hugo-Coetzee, E.A. 2014. New Oppiidae (Acari: Oribatida) from the Golden Gate Highlands National Park in South Africa. Zootaxa 3884(6): 533-552.

Coetzee, L. 2013. New species of the genus Afroleius Mahunka, 1984 (Acari, Oribatida, Mycobatidae) from South Africa. Acta Zoologica Academiae Scientiarum Hungaricae 59(4): 307-319. PDF 1.9 MEG

Coetzee, L. & Tiedt, L.R. 2013. Overview of the genus Afroleius Mahunka, 1984 Acari, Oribatida). In: Schausberger, P. (ed.) Acari in a changing world: Proceedings of the 7th Symposium of EURAAC, Vienna, 2012. Acarologia 53(2):163-173. PDF 1.1MEG

Hugo-Coetzee, E.A. 2013. New species of Aleurodamaeus Grandjean, 1954 (Oribatida: Aleurodamaeidae) from South Africa. Zootaxa 3670(4): 531-556.

Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species of Nothrus (Acari: Oribatida: Nothridae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 28(2): 25-40. PDF 850 KB

Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Valbehanella freestatensis gen. nov., sp. nov. (Acari, Oribatida) from South Africa.  Systematic & Applied Acarology 17(4): 407-416.

Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Oribatid mites of the genus Eremulus Berlese, 1908 (Acari: Oribatida: Eremulidae) from South Africa.  African Invertebrates 53(2): 559-569. PDF 860KB

Schuster, R.K. & Coetzee, L. 2012. Cysticercoids of Anoplocephala magna (Eucestoda: Anoplocephalidae) experimentally grown in oribatid mites (Acari: Oribatida). Veterinary Parasitology. view link

Coetzee, L. & Weigmann, G. 2012. Systematic revision of the genus Maudheimia Dalenius, 1958 (Acari: Oribatida). Zootaxa 3295: 65-68.

Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. The oribatid mite genus Nothrus Koch, 1836 (Acari: Oribatida: Nothridae) of South Africa, including a key to African species.  Zootaxa 3243: 29-51.

Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species from South Africa, with remarks on generic diagnosis of Licnodamaeolus Covarrubias, 1998 and taxonomic status of Nacunansella Fernandez & Cleva, 1998 (Acari: Oribatida: Licnodamaeidae). Zootaxa 3167: 32-44.

Hugo-Coetzee, E.A. & Avenant, N.L. 2011. The effect of fire on soil oribatid mites (Acari: Oribatida) in a South African grassland.  Zoosymposia 6: 210-220.

Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov, A.A. 2011. Morphology of juvenile instars of Neoliodes terrestris (Wallwork, 1963) and N. ionicus Sellnick, 1931 (Acari: Oribatida: Neoliodidae).  Annales Zoologici (Warszawa) 61(4): 817-830.

Hugo-Coetzee, E.A. 2011.  Three new species of Austrocarabodes (Oribatida: Carabodidae) and notes on Austrocarabodes pinnatus Mahunka, 1986, from South Africa.  Zootaxa 3011: 1-15.

Hugo-Coetzee, E.A. 2010. Two new species of Austrocarabodes (Acari: Oribatida: Carabodidae) from South Africa: A. mahunkai and A. lineasetosa.  Navorsinge van die Nasionale Museum, Bloemfontein26(2): 45-58. PDF 1 MB

Hugo, E.A. 2010. Two new species of Gymnodamaeidae (Acari: Oribatida) from South Africa.  International Journal of Acarology  36(3): 199-210.

Coetzee, L. 2010. Species or morphological variation? A multivariate morphometric analysis of Afroleius simplex (Acari, Oribatida, Haplozetidae). M.W. Sabelis & J. Bruin (eds).  Trends in Acarology.  pp. 267-269.  PDF 926KB

Sinclair, W., Coetzee, L. & Joubert, G. 2010. House-dust mite species in Bloemfontein, South Africa. South African Medical Journal 95(3): 630-634. PDF 340KB

Hugo, E.A. 2008. Three new species of Austrocarabodes (Oribatida: Carabodidae) from South Africa. Zootaxa 1844: 25-36.

Coetzee, L. 2007.  The genus Afroleius Mahunka (Acari, Oribatida, Haplozetidae) 1. Redescriptions of A. deformis, A. minor and A. simplex. Navorsinge van die Nasionale Museum, Bloemfontein 23(3): 101-118.  PDF 1,6MEG.

Akrami, M.A. & Coetzee, L. 2007. Mabulatrichus iranicus (Acari: Oribatida: Zetomotrichidae): a new species from Iran. Systematic and Applied Acarology 12: 245-252.  PDF 93KB

Hugo, E.A., Chown, S.L. & McGeoch, M.A. 2006. The microarthropods of sub-Antarctic Prince Edward Island: a quantitative assessment. Polar Biology, 30(1):109-119  PDF 208KB

Grobler, L., Bayram, S. & Cobanoglu, S. 2005. Two new records of Oribatula (Zygoribatula) species (Acari: Oribatida) from Turkey, with redescriptions. Zoological Science 22:1347-1351.

Hugo, E.A., McGeoch, M.A., Marshall, D.J. & Chown, S.L. 2004. Fine scale variation in microarthropod communities inhabiting the keystone species Azorella selago on Marion Island. Polar Biology, 27(8):466-473.  PDF 201KB

Yamamoto, Y. & Coetzee, L. 2004. The Trimalaconothrus species from South Africa (Acari: Oribatida: Malaconothridae). Acta Arachnologica 53(1): 13-25.  PDF 3.6 MEG

Grobler, L., Bayram, S., & Cobanoglu, S. 2004. Two new species and new records of Oribatid mites from Turkey. International Journal of Acarology 30(4): 351-358.

Yamamoto, Y. & Coetzee, L. 2003. The Malaconothrus species from South Africa and Lesotho (Acari: Oribatida: Malaconothridae). Acta Arachnologica 52(2): 113-126. PDF 3.4 MEG

Grobler, L., Ozman, S.K. & Cobanoglu, S. 2003. The genera Liacarus, Stenoxenillus and Xenillus (Oribatida: Gustavioidea) from Turkey. Acarologia 43(1): 133-149.

Grobler, L. 2003. The genus Furcoppia (Acari, Oribatida) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 19(7): 161-179

Coetzee, L. & Brink, J.S. 2003. Fossil oribatid mites (Acari, Oribatida) from the Florisbad Quaternary deposits, South Africa. Quaternary Research 59: 246-254.  PDF 3,3MEG

Coetzee, L. & Marshall, D.J., 2003. A new Halozetes species (Acari: Oribatida: Ameronothridae) from the marine littoral of southern Africa. African Zoology 38(2): 327-331.  PDF 427KB

Coetzee, L. 2003. A new genus and species Floritrichus louisbothai (Acari, Oribatida, Oripodoidea, Zetomotrichidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 9(5):93-100. PDF 335KB

Coetzee, L. 2002. Preliminary report on fossil oribatid mites from the Florisbad quaternary research station, South Africa. F. Bernini et al. (eds). Acarid Phylogeny an Evolution Adaptations in mites and ticks, pp. 41-44. PDF 1,3 MEG

Coetzee, L. 2001. Lohmanniidae species (Acari, Oribatida) from the Holocene Deposits at Florisbad, South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 17(5):125-134.  PDF 237KB

Coetzee, L. 2001. New species of the Family Lohmanniidae (Acari, Oribatida) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 17(3):53-67. PDF 327KB

Heyne, H., Ueckermann, E.A. & Coetzee. L. 2001. First report of a parasitic mite, Leptotrombidium subquadratum (Lawrence) (Acari: Trombiculidae, Trombiculinae), from dogs and children in the Bloemfontein area, South Africa. Journal of the South African Veterinary Association. 72(2):105-106. PDF 1,3 MEG

Coetzee, L. 2000. A new subspecies of Halozetes belgicae (Acari, Oribatida, Ameronothridae) from the Marine supra-littoral of the sub-Antarctic Marion Island. Acarologia. 15(4):439-447.  PDF 2MEG

Grobler, L & Skubala, P. 2000. Description of Oribatula dentata, sp. nov. (Oribatulidae, Oribatida) from Spain and complementary data on O. longolamellata and O. macrostega. Acarologia 15(3): 343-353.

Marshall, D.J. & Coetzee, L. 2000. Historical biogeography and ecology of a Continental Antarctic mite genus, Maudheimia (Acari, Oribatida): evidence for a Gondwanan origin and Pliocene-Pleistocene speciation. Zoological Journal of the Linnean Society 129:111-128. PDF 2,6 MEG

Bayartogtokh, B., Grobler, L. & Cobanoglu, S. 2000. A new species of Punctoribates (Acari: Oribatida: Mycobatidae) collected from mushrooms in Turkey, with remarks on the taxonomy of the genus. Navorsinge van die Nasionale Museum, Bloemfontein 16(2):18-32.

Schuster, R., Coetzee. L. & Putterill, J.F. 2000. Oribatid mites (Acari, Oribatida) as intermediate hosts of tapeworms of the Family Anoplocephalidae (Cestoda) and the transmission of Moniezia expansa cysticercoids in South Africa. Onderstepoort Journal of Veterinary Research 67:49-55.  PDF 1.4 MEG

Grobler, L. 2000. Megatrichobates Gen. Nov. (Acari, Oribatida, Oribatulidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 16(1):1-14.

Marshall, D.J., Gremmen, N.J.M., Coetzee, L., O'Connor, B.M., Pugh, P.J.A., Theron, P.D. & Ueckermann, E.A. 1999. New records of Acari from the sub-Antarctic Prince Edward Islands. Polar Biology 21:84-89.  PDF 2,0 MEG

Grobler, L. 1998. Subfamily Pseudotocepheinae (Acari, Oribatida, Otocepheidae): Constrictocepheusgen. nov. and new Pseudotocepheus species from South Africa Navorsinge van die Nasionale Museum, Bloemfontein 14(1): 1-44.

Coetzee, L. 1997. The Antarctic mite genus Maudheimia (Acari, Oribatida). Navorsinge van die Nasionale Museum, Bloemfontein 13 (3): 101-135.  PDF 4,2 MEG

Grobler, L. 1997. The Subfamily Pseudotocepheinae subfam. nov. (Acari, Oribatida, Otocepheidae). Navorsinge van die Nasionale Museum, Bloemfontein 13(1): 1-44.

Grobler, L. 1995. Two Otocepheid genera (Acari, Oribatida) from South Africa: Longocepheus Balogh & Mahunka and Trichocepheus Balogh & Mahunka. Navorsinge van die Nasionale Museum, Bloemfontein 11(10): 261-300.

Grobler, L. 1995. New species of the genus Plenotocepheus Hammer, 1966 (Acari, Oribatida, Otocepheidae) from South Africa II. Navorsinge van die Nasionale Museum,  Bloemfontein 11(5): 102-118.

Grobler, L. 1995. New species of the genus Plenotocepheus Hammer, 1966 (Acari, Oribatida, Otocepheidae) from South Africa I. Navorsinge van die Nasionale Museum,  Bloemfontein 11(2): 13-36.

Coetzee, L. 1995. A new species of the genus Saltatrichus Coetzee, 1993 (Acari, Oribatida, Zetomotrichidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 11(1): 1-12.  PDF 1,1 MEG

Grobler, L. 1994. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa III. Zygoribatula contracta spec. nov. and Zygoribatula setosa Evans, 1953. Navorsinge van die Nasionale Museum, Bloemfontein 10(6): 261-279.

De Wet, L. 1993. New species of the genus Hermannia Nicolet, 1855 from South Africa II. (Acari, Oribatida, Hermanniidae). Navorsinge van die Nasionale Museum, Bloemfontein 9(9): 266-287.

Grobler, L. & Kok, D.J. 1993. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa II. New and existing species. Navorsinge van die Nasionale Museum, Bloemfontein 9(6): 182-212.

Coetzee, L. 1993. New genera and species of the family Zetomotrichidae Grandjean, 1954 (Acari, Oribatida, Oripodoidea) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 9(5): 133-179.  PDF 4,1 MEG

Grobler, L. 1993. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa I. New species. Navorsinge van die Nasionale Museum,  Bloemfontein 9(3): 49-76.

De Wet, L. 1993. New species of the genus Hermannia Nicolet, 1855 from South Africa I. (Acari: Oribatida: Hermanniidae). Navorsinge van die Nasionale Museum, Bloemfontein 9(2): 21-48.

Grobler, L. 1991. A new Brasilobates Perez-Inigo & Baggio, 1980 from South Africa (Acari: Oribatei: Oribatuloidea: Xylobatidae). Navorsinge van die Nasionale Museum,  Bloemfontein 7(5): 73-95.

Grobler, L. 1989. New South African species of the genus Eupelops Ewing, 1917 (Acari: Oribatei: Phenopelopoidea: Phenopelopidae). Navorsinge van die Nasionale Museum,  Bloemfontein 6(5): 151-201.

Coetzee, L. 1989. The genus Zetorchestes Berlese, 1888 (Acari, Oribatei, Zetorchestidae) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 6(6): 203-221.  PDF 1,5 MEG

Coetzee, L. 1987. The South African Lamellareidae Balogh, 1972 (Acari: Oribatei). Navorsinge van die Nasionale Museum, Bloemfontein 5(13): 325-353.  PDF 2,9 MEG

Engelbrecht, C.M. 1986. New Oribatid taxa and distribution records predominantly from Southern Africa. (Acari: Oribatei: Oribatelloidea). Navorsinge van die Nasionale Museum, Bloemfontein 5(9): 170-250.

Engelbrecht, C.M. 1975. Die Suid-Afrikaanse Eupelopoidea Balogh, 1963. Navorsinge van die Nasionale Museum, Bloemfontein 3(5): 89-108.

Engelbrecht, C.M. 1975. New Ameronothroid (Oribatei, Acari) taxa from the Republic of South Africa and the Islands Gough and Marion. Navorsinge van die Nasionale Museum,  Bloemfontein 3(4): 53-88.

Engelbrecht, C.M. 1974. The genus Hydrozetes Berlese, 1902. (Oribatei: Acari) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 3(3): 41-49.

Engelbrecht, C.M. 1974. The genus Passalozetes Grandjean, 1932 (Oribatei: Acari) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 3(2): 29-38.

Engelbrecht, C.M. 1974. The genus Halozetes (Oribatei: Acari) on Marion Island. Navorsinge van die Nasionale Museum, Bloemfontein 3(1): 1-25.

Engelbrecht, C.M. 1973. Magoebazetes, a new microzetid genus (Microzetidae: Oribatei) from South Africa. Acarologia 14(4): 681-695.

Engelbrecht, C.M. 1973. South African Galumnoidea (Oribatei: Acari): New taxa and records. Navorsinge van die Nasionale Museum, Bloemfontein 2(11): 394-415.

Engelbrecht, C.M. 1972. A new South African microzetid genus, Afrozetes (Microzetidae: Oribatei). Acarologia 14(3): 484-496.

Engelbrecht, C.M. 1972. Ctenogalumna moresonensis sp. n. and Pilogalumna bloemfonteinensis sp. n., two new South African species of the subfamily Allogalumninae Balogh, 1960 (Galumnidae: Oribatei). Acarologia 14(3): 497-510.

Engelbrecht, C.M. 1972. South African species of the genus Trichogalumna Balogh, 1961 (Galumnidae: Oribatei). Acarologia 14(2): 268-280.

Engelbrecht, C.M. 1972. Galumnids from South Africa. Acarologia 14(1): 109-140.

Engelbrecht, C.M. 1972. Dimidiogalumna  villiersensis (Galumnidae: Oribatei), a new South African galumnid genus and its type species. Acarologia 14(1): 141-149.

Engelbrecht, C.M. 1972. A new microzetid genus and species Schalleriella grabouwensis(Microzetidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum,  Bloemfontein 2(9): 282-285.

Engelbrecht, C.M. 1972. Microzetes auxiliaris orkneyensis s. sp. n. and Microzetes natalensis sp. n. (Microzetidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 270-281.

Engelbrecht, C.M. 1972. Two new species of the genus Rhopalozetes Balogh, 1961 Rhopalozetes georgensis sp. n. and Rhopalozetes witfonteinensis sp. n. (Microzetidae: Oribatei). Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 263-269.

Engelbrecht, C.M. 1972. Two new species of the genus Galumna Von Heyden, 1826 (Galumnidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 244-262.

Engelbrecht, C.M. 1969. Some South African species of the genus Galumna Von Heyden, 1826 (Acari: Galumnidae). Journal of the Entomological Society of Southern Africa 32(1): 99-122.

Articles from Culna 60 onwards available at culna-downloads/

Hugo-Coetzee, L. 2016. Mites at the beach. Culna 71: 14-15

Coetzee, L. 2015. Die alomteenwoordige myte. Culna 70: 20-21.

Hugo-Coetzee, L. 2014. Soil mites on guard. Culna 69:24-25.

Coetzee, L. 2013. Grondmyte en die stryd teen waterhiasinte. Culna 68:11-13.

Hugo-Coetzee, L. 2013. What's in a name? From Hitler to the vuvuzela. Culna 68: 20-21.

Hugo-Coetzee, L. 2012. Die kortafstand ... vir insekte. Culna 67: 26-27

Coetzee, L. 2011. Myte - het ons hulle nodig? Culna 66: 12-14

Hugo-Coetzee, L. 2011. Living Soil. Culna 66: 22-23

Coetzee, L. 2009. Ongewone bedmaats - huisstofmyte. Culna 64: 29-39

Hugo, L. 2009. Describing species - how it is done.  Culna 64: 19-20

Hugo, L. 2008. Goggas in die water. Culna 63: 17-18

Hugo, L. 2007. The sex life of soil mites. Culna 62: 16-17

Coetzee, L. 2006. The Secretarybird - majestic hunter of the plains. Culna 61: 30-31

Hugo, L. 2006. Myte - die agtbenige "ysbere" van Antarktika. Culna 61: 16-17

Coetzee, L. 2005. Chiggers - are your dogs biting their paws? Culna 60: 19-20

Coetzee, L. 2004. What makes an owl? Culna 59: 17-19

Coetzee, L. 2000. Oribatid mites found in the Quaternary sediments at Florisbad. Culna 55: 21

Coetzee, L. 1998. Wat gaan binne in die grond aan? Culna 53: 16

Coetzee, L. 1984. Oribatei. National Museum News 27: 26-27

Taxonomic research is focussed mainly on the Oribatida.

The collection currently consists of 27 220 oribatid samples (each containing a varying number of specimens), collected from 4 580 localities.  The samples were collected from a wide range of habitats in South Africa, Gough Island and the sub Antarctic Islands Marion and Prince Edward. The specimens are stored in 70% Alcohol + 5% Glycerol.

The holotype collection consists of 223 Oribatida, 19 Astigmata (Analgoidea), collected from birds in South Africa, and 13 Prostigmata (Hygrobatoidea).

The collection of Oribatida holotypes which belongs to the Ditsong National Museum of Natural History, Pretoria,  (formerly the Transvaal Museum), are now at the National Museum, Bloemfontein, on a long term loan. This material comprises oribatids collected from South Africa and studied by Dr. Sandor Mahunka.

Astigmata Holotypes

• Analgidae
  

  Analgidae

  Anagles vanasi Mironov & Kopij, 1996

• Proctophyllodidae
  

  Proctophyllodidae	Montesauria belozerovi Mironov & Kopij, 1996
  	Montesauria emberizae Mironov & Kopij, 1996
  	Montesauria jesionowskii Mironov & Kopij, 1997
  	Montesauria punctata Mironov & Kopij, 1996
  	Montesauria tetralobula Mironov & Kopij, 1996
  	Pedanodectes marginatus Mironov & Kopij, 1997
  	Proctophyllodes caffer Mironov & Kopij, 1996
  	Proctophyllodes leptodiscus Mironov & Kopij, 1996

• Pteronyssidae
  
  

  Pteronyssidae	Metapteronyssus angolensis Mironov & Kopij, 2000
  	Pteroherpus africanus Mironov & Kopij, 2000
  	Pteronyssoides nectariniae Mironov, 2001
  	Pteronyssoides promeropis Mironov & Kopij, 2000
  	Sturnotrogus creatophorae Mironov & Kopij, 2000

• Trouessartiidae
  
  

  Trouessartiidae	Trouessartia angolensis Mironov & Kopij, 2000
  	Trouessartia creatophorae Mironov & Kopij, 1996
  	Trouessartia daberti Mironov & Kopij, 2000
  	Trouessartia priniae Mironov & Kopij, 2000
  	Trouessartia pytiliae Mironov & Kopij, 2000

Oribatida Holotypes

• Achipteriidae
  

  Achipteriidae	Novoribatella minutisetarum Engelbrecht, 1986
  	Novoribatella pentasetarum Engelbrecht, 1986

• Aleurodamaeidae
  

  Aleurodamaeidae	Aleurodamaeus salvadordalii Hugo-Coetzee, 2013
  	Aleurodamaeus vicinus Hugo-Coetzee, 2013
  	Aleurodamaeus angelae Hugo-Coetzee, 2013
  	Aleurodamaeus niedbalai Hugo-Coetzee, 2013
  	Aleurodamaeus minutus Hugo-Coetzee, 2013
  	Aleurodamaeus woasi Hugo-Coetzee, 2013
  	Aleurodamaeus prominens Hugo-Coetzee, 2013

• Ameronothridae
  

  Ameronothridae	Alaskozetes bouvetoyaensis  Van Pletzen & Kok
  	Halozetes belgicae mickii  Coetzee, 2000
  	Halozetes edwardensis  Van Pletzen & Kok
  	Halozetes fulvus  Engelbrecht, 1975
  	Halozetes marinus devilliersi  Engelbrecht, 1974
  	Halozetes marionensis  Engelbrecht, 1974

• Aquanothridae
  

  Aquanothridae

  Aquanothrus montanus Engelbrecht, 1975

• Carabodidae
  

  Carabodidae	Austrocarabodes hendriksi Hugo, 2008
  	Austrocarabodes luciensis Hugo, 2008
  	Austrocarabodes vernoncrookensis Hugo, 2008
  	Austrocarabodes mahunkai Hugo-Coetzee, 2010
  	Austrocarabodes lineasetosa Hugo-Coetzee, 2010
  	Austrocarabodes nortoni Hugo-Coetzee, 2011
  	Austrocarabodes longisetosus Hugo-Coetzee, 2011
  	Austrocarabodes crassimarginatus Hugo-Coetzee, 2011

• Crotonidae
  

  Crotonidae	Crotonia dicella Colloff, 1990
  	Crotonia ecphyla Colloff, 1990
  	Crotonia lyrata Colloff, 1990
  	Crotonia pauropelor Colloff, 1990

• Cymbaeremaeidae
  

  Cymbaeremaeidae	Scapheremaeus pundamiliaensis  Engelbrecht, 1975
  	Scapheremaeus rustenburgenesis  Engelbrecht, 1975

• Eremaeozetidae
  

  Eremaeozetidae	Eremaeozetes capensis Schatz, 2001
  	Eremaeozetes louisae Schatz, 2003
  	Eremaeozetes nasutus Schatz,  2003
  	Mahunkaia tricornis Schatz, 2002

• Eremulidae
  

  Eremulidae	Eremulus spindleformis Ermilov & Hugo-Coetzee, 2012
  	Eremulus southafricanensis Ermilov & Hugo-Coetzee, 2012

• Euphthiracaridae
  

  Euphthiracaridae	Acrotritia minuta Niedbała, 2006
  	Euphthiracarus rectus Niedbała, 2006
  	Euphthiracarus scuticus Niedbała, 2006

• Galumnellidae
  

  Galumnellidae

  Galumnella pilosa Engelbrecht, 1972

• Galumnidae
  

  Galumnidae	Ctenogalumna moresonensis Engelbrecht, 1972
  	Dimidiogalumna villiersensis Engelbrecht, 1972
  	Galumna capensis capensis Engelbrecht, 1969
  	Galumna capensis dissimilis Engelbrecht, 1969
  	Galumna comparabilis Engelbrecht, 1969
  	Galumna dimidiata Engelbrecht, 1969
  	Galumna nuda Engelbrecht, 1969
  	Galumna rostrata Engelbrecht, 1969
  	Galumna tricuspidata Engelbrecht, 1969
  	Pergalumna elongata Engelbrecht, 1972
  	Pergalumna incomperta Engelbrecht, 1972
  	Pergalumna magnipora capensis Engelbrecht, 1972
  	Pilogalumna bloemfonteinensis  Engelbrecht, 1972
  	Pilogalumna kimberleyensis Engelbrecht, 1972
  	Pilogalumna variabilis Engelbrecht, 1972
  	Sacculogalumna saccularis Engelbrecht1973
  	Trichogalumna pumularis Engelbrecht, 1972
  	Trichogalumna punctata Engelbrecht, 1972

• Gymnodamaeidae
  

  Gymnodamaeidae	Arthrodamaeus johanni Hugo, 2009
  	Gymnodamaeus deswardti Hugo, 2009
  	Adrodamaeus ermilovi Hugo-Coetzee, 2014

• Hermanniidae
  

  Hermanniidae	Hermannia comparabilis De Wet, 1993
  	Hermannia engelbrechti De Wet, 1993
  	Hermannia exobothridialis De Wet, 1993
  	Hermannia louisiae De Wet, 1993
  	Hermannia natalensis De Wet, 1993
  	Hermannia nathanaeli De Wet, 1993
  	Hermannia notogastralis De Wet, 1993
  	Hermannia phylliformis De Wet, 1993
  	Hermannia setiformis De Wet, 1993
  	Hermannia transvaalensis De Wet, 993

• Hydrozetidae
  

  Hydrozetidae

  Hydrozetes capensis Engelbrecht, 1975

• Lamellareidae
  

  Lamellareidae	Lamellarea ardua Kok, 1968
  	Lamellarea digitata Kok, 1968
  	Lamellarea forceps Kok, 1968
  	Lamellarea quadrata Coetzee, 1986
  	Microlamellarea engelbrechti Coetzee, 1986

• Licnodamaeidae
  

  Licnodamaeidae	Licnodamaeolus asetosus Ermilov & Hugo-Coetzee, 2012
  	Licnodamaeolus erfenisdamensis Ermilov & Hugo-Coetzee, 2012

• Lohmanniidae
  

  Lohmanniidae	Annectacarus eksteeni Coetzee, 2001
  	Papillacarus brinki Coetzee, 2001
  	Paulianacarus barlowi Coetzee, 2001
  	Paulianacarus grobleri Coetzee, 2001

• Malaconothridae
  

  Malaconothridae	Malaconothrus longidorsus Yamamoto & Coetzee, 2004
  	Malaconothrus minimus Yamamoto & Coetzee, 2004
  	Malaconothrus stigmatus Yamamoto & Coetzee, 2004
  	Trimalaconothrus binodulus Yamamoto & Coetzee, 2004
  	Trimalaconothrus duaculeus Yamamoto & Coetzee, 2004
  	Trimalaconothrus obesus Yamamoto & Coetzee, 2004
  	Trimalaconothrus punctum Yamamoto & Coetzee, 2004
  	Trimalaconothrus rectus Yamamoto & Coetzee, 2004

• Maudheimiidae
  

  Maudheimiidae	Maudheimia marshalli  Coetzee, 1997
  	Maudheimia tanngardenensis  Coetzee, 1997

• Mesoplophoridae
  

  Mesoplophoridae	Mesoplophora iuvenalis Niedbała, 2006
  	Mesoplophora pertenuis Niedbała, 2006
  	Mesoplophora setulosa Niedbała, 2006

• Metrioppiidae
  

  Metrioppiidae	Furcoppia dentata Grobler, 2003
  	Furcoppia longiseta Grobler, 2003
  	Furcoppia minuta Grobler, 2003

• Microzetidae
  

  Microzetidae	Afrozetes champaignensis  Engelbrecht, 1972
  	Afrozetes magoebaensis capensis  Engelbrecht, 1972
  	Afrozetes magoebaensis magoebaensis  Engelbrecht, 1972
  	Afrozetes natalensis  Engelbrecht, 1972
  	Magoebazetes magoebaensis  Engelbrecht, 1973
  	Magoebazetes natalensis  Engelbrecht, 1973
  	Magoebazetes pentasetarum  Engelbrecht, 1973
  	Microzetes auxiliaris orkneyensis  Engelbrecht, 1972
  	Microzetes natalensis  Engelbrecht, 1972
  	Rhopalozetes georgensis  Engelbrecht, 1972
  	Rhopalozetes witfonteinensis  Engelbrecht, 1972
  	Schalleriella grabouwensis  Engelbrecht, 1972

• Nothridae
  

  Nothridae	Nothrus bilongisetosus Ermilov & Hugo-Coetzee, 2012
  	Nothrus monolongisetosus Ermilov & Hugo-Coetzee, 2012
  	Nothrus louiseae Ermilov & Hugo-Coetzee, 2012
  	Nothrus engelbrechti Ermilov & Hugo-Coetzee, 2012

• Oppiidae
  

  Oppiidae	Amerioppia africana Kok, 1967
  	Brachioppia longisetosa Kok, 1967
  	Brachioppia moresonensis Kok, 1967
  	Brachioppia orkneyensis Kok, 1967
  	Brachioppia pectinata Kok, 1967
  	Brachioppia quathlambae Kok, 1967
  	Oppia falxa Kok, 1967
  	Oppia parva Kok, 1967
  	Oppia pletzenae Kok, 1967
  	Oppia quattuor Kok, 1967
  	Oppia yodai africana Kok, 1967
  	Brachioppia louwi Hugo-Coetzee, 2014
  	Brachioppiella (B) dawidi Hugo-Coetzee, 2014
  	Brachioppiella (B) goblina Hugo-Coetzee, 2014
  	Kokoppia mandelai Hugo-Coetzee,  2014
  	Lanceoppia scytheae Hugo-Coetzee, 2014
  	Oppiella (Quattoppiella) goldengatensis Hugo-Coetzee, 2014

• Oribatulidae
  

  Oribatulidae	Megatrichobates longisetosus Grobler, 2000
  	Megatrichobates striatus Grobler, 2000
  	Zygoribatula connexa substriata Grobler, 1993
  	Zygoribatula gracilata Grobler, 1993
  	Zygoribatula lineaporosa Grobler, 1993
  	Zygoribatula pennata Grobler, 1993
  	Zygoribatula robusta Grobler, 1993
  	Zygoribatula subantarctica Van Pletzen & Kok

• Oribotritiidae
  

  Oribotritiidae	Austrotritia engelbrechti Niedbała, 2006
  	Indotritia cypha Niedbała, 2006
  	Indotritia didyma Niedbała, 2006
  	Indotritia eksteeni Niedbała, 2006
  	Indotritia fusa Niedbała, 2006
  	Indotritia partita Niedbała, 2006
  	Indotritia phymatha Niedbała, 2006
  	Oribotritia deminuta Niedbała, 2006

• Otocepheidae
  

  Otocepheidae	Longocepheus globosus Grobler, 1995
  	Plenotocepheus crinitus Grobler, 1995
  	Plenotocepheus dentatus Grobler, 1995
  	Plenotocepheus discrepans Grobler, 1995
  	Plenotocepheus verrucosus Grobler, 1995
  	Pseudotocepheus aokii Grobler, 997
  	Pseudotocepheus contractus Grobler, 1997
  	Pseudotocepheus dentatus Grobler, 1998
  	Pseudotocepheus lanceolatus Grobler, 1998
  	Pseudotocepheus promissus Grobler, 1998
  	Pseudotocepheus shauni Grobler, 1997
  	Trichocepheus auriculus Grobler, 1995

• Parakalummidae
  

  Parakalummidae

  Carinogalumna montana Engelbrecht, 1973

• Passalozetidae
  

  Passalozetidae	Passalozetes douglasensis  Engelbrecht, 1974
  	Passalozetes wolwekransensis  Engelbrecht, 1974

• Phenopelopidae
  

  Phenopelopidae	Eupelops capensis  Grobler, 1989
  	Eupelops curtipilus thabazimbiensis  Engelbrecht, 1975
  	Eupelops distinctus  Grobler, 1989
  	Eupelops engelbrechti  Grobler, 1989
  	Eupelops foveolatus  Engelbrecht, 1975
  	Eupelops margatensis  Engelbrecht, 1975
  	Eupelops minutus  Grobler, 1989

• Phthiracaridae
  

  Phthiracaridae	Phthiracarus anakolos Niedbała, 2006
  	Phthiracarus blythedalensis Niedbała, 2006
  	Phthiracarus densus Niedbała, 2006
  	Phthiracarus hillcrestensis Niedbała, 2006
  	Phthiracarus humilis Niedbała, 2006
  	Phthiracarus kokae Niedbała, 2006
  	Phthiracarus leliehoekensis Niedbała, 2006
  	Phthiracarus pertenuis Niedbała, 2006

• Punctoribatidae
  

  Punctoribatidae	Afroleius crassus Coetzee, 2013
  	Afroleius decurvatus Coetzee, 2013
  	Afroleius deformatoides Coetzee, 2013
  	Afroleius amieae Coetzee, 2014
  	Afroleius inae Coetzee, 2014
  	Afroleius valerieae Coetzee, 2014
  	Afroleius natalensis Coetzee, 2014
  	Afroleius caudatus Coetzee, 2014
  	Afroleius lucidus Coetzee, 2015

• Scheloribatidae
  

  Scheloribatidae	Scheloribates durbanensis Van Pletzen, 1963
  	Scheloribates elsi Van Pletzen, 1965
  	Scheloribates louwi Van Pletzen, 1965
  	Scheloribates minutus Van Pletzen, 1965
  	Scheloribates natalensis Van Pletzen, 1963
  	Scheloribates parvus Van Pletzen, 1963
  	Scheloribates robustus Van Pletzen, 1963
  	Scheloribates rustenburgensis Van Pletzen, 1963
  	Scheloribates tzitzikamaenesis Van Pletzen, 1965
  	Scheloribates variabilis Van Pletzen, 1965

• Scutoverticidae
  

  Scutoverticidae	Scutovertex sculperens Kok, 1968
  	Scutoverticosus arcanus Kok, 1968

• Steganacaridae
  

  Steganacaridae	Arphthicarus bulbosus  Niedbała, 2006
  	Atropacarus brevisetosus  Niedbała, 2006
  	Austrophthiracarus cordylus  Niedbała, 2006
  	Notophthiracarus agulhasensis  Niedbała, 2006
  	Notophthiracarus capevidalensis  Niedbała, 2006
  	Notophthiracarus coetzeeae  Niedbała, 2006
  	Notophthiracarus cristatus  Niedbała, 2006
  	Notophthiracarus frondeus  Niedbała, 2006
  	Notophthiracarus gongylos  Niedbała, 2006
  	Notophthiracarus grossus  Niedbała, 2006
  	Notophthiracarus knysnaensis  Niedbała, 2006
  	Notophthiracarus korannabergensis  Niedbała, 2006
  	Notophthiracarus mekistos  Niedbała, 2006
  	Notophthiracarus natalensis  Niedbała, 2006
  	Notophthiracarus phyllodes  Niedbała, 2006
  	Notophthiracarus procerus  Niedbała, 2006
  	Notophthiracarus pumilus  Niedbała, 2006
  	Notophthiracarus rhachis  Niedbała, 2006
  	Notophthiracarus serratus  Niedbała, 2006
  	Notophthiracarus tsitsikamaensis  Niedbała, 2006
  	Notophthiracarus vernonensis  Niedbała, 2006
  	Notophthiracarus zululandensis  Niedbała, 2006
  	Protophthiracarus crinatus  Niedbała, 2006
  	Protophthiracarus engelbrechti  Niedbała, 2006

• Tectocepheidae
  

  Tectocepheidae	Tegeocranellus knysnaensis Kok, 1968
  	Tegeocranellus sacchareus Kok, 1968

• Tegoribatidae
  

  Tegoribatidae	Tegoribates montana Engelbrecht, 1986
  	Tegoribates natalensis Engelbrecht, 1986
  	Tegoribates nuda Engelbrecht, 1986

• Unduloribatidae
  

  Unduloribatidae

  Valbehanella freestatensis Ermilov & Hugo-Coetzee, 2012

• Xylobatidae
  

  Xylobatidae

  Brasilobates punctata Grobler, 1991

• Zetomotrichidae
  

  Zetomotrichidae	Demisalto engelbrechti Coetzee, 1993
  	Floritrichus louisbothai Coetzee, 2003
  	Mabulatrichus dentatus Coetzee, 1993
  	Saltatrichus magnus Coetzee, 1993
  	Saltatrichus marshalli Coetzee, 1995
  	Saltatrichus minus Coetzee, 1993
  	Saltatrichus robustus Coetzee, 1993

• Zetorchestidae
  

  Zetorchestidae

  Zetorchestes transvaalensis Coetzee, 1989

Prostigmata Holotypes

• Hygrobatidae
  

  Hygrobatidae	Hygrobates draconicus Smit, 2005
  	Hygrobates inflatipes Smit, 2005
  	Hygrobates laceratoides Smit, 2005
  	Hygrobates spathuliferus cederberg Smit, 2005
  	Atractides capensis Pesic, Smit & Gerecke, 2011
  	Atractides cooki Pesic, Smit & Gerecke, 2011
  	Atractides dracomontanus  Pesic, Smit & Gerecke, 2011
  	Atractides meridianus  Pesic, Smit & Gerecke, 2011
  	Atractides neoscutifer Pesic, Smit & Gerecke, 2011

• Torrenticolidae
  

  Torrenticolidae	Monatractides nigromontanus  Goldschmidt & Smit, 2009
  	Monatractides drakenbergensis  Goldschmidt & Smit, 2009
  	Torrenticolidae spatulifra  Goldschmidt & Smit, 2009
  	Torrenticola tsitsikamma  Goldschmidt & Smit, 2009

The acceptance of material on loan implies that the following conditions have been accepted:

1. Loan material must be handled with the utmost care and must be securely packed for return by insured or registered mail.
2. National Museum specimens described as holotypes must be returned.
3. All labels and numbers given to specimens/objects must be retained and under no circumstances should any of these be discarded or altered. Where applicable the catalogue numbers of specimens/objects must be given in descriptions.
4. The National Museum should be credited in any publication when referring to this material.