The Study of Mites
The Acarology Department (study of mites) came into existence as an independent department at the National Museum in 1969. Prof. R. van Pletzen, former Head of Zoology at the University of the Free State, started oribatid mite taxonomy research at the university. One of his students, Dr C.M. Engelbrecht, took up a post at the National Museum in 1969 and continued the research at the Museum. The research focus at the University changed and the collection of specimens and literature was donated to the Museum. Dr Engelbrecht collected widely throughout South Africa, augmenting the collection fourfold. Research is mainly focussed on taxonomy of Oribatida.
WHAT ARE ORIBATID MITES?
Oribatid mites, also known as soil mites or moss mites, belong to the Class Arachnida, Subclass Acari, Suborder Oribatida (=Cryptostigmata). They range in size from 0.3 – 0.7 mm (with extremes from 0.15 – 2 mm) and occur mainly in the top layer of soil and plant litter, where they occur in large numbers. Abundance and diversity of oribatids vary greatly according to soil conditions. Certain species are also adapted to live on plants, in mosses and lichens and in the marine littoral zone. Soil mites are mostly particle-feeding saprophages (feeding on decaying organic matter) and mycophages (fungus-eating), thereby playing an important role in decomposition of organic material, soil fertility, nutrient cycling and soil formation.
Acarologists appointed Research Associates
11 January 2017
Drs Louise Coetzee (left) and Lizel Hugo-Coetzee have been appointed Research Associates in the Faculty of Natural and Agricultural Sciences at the Univeristy of the Free State. They will be collaborating with the Department of Zoology and Entomology.
EURAAC in Valencia, Spain
10 January 2017
Dr Lizel Hugo-Coetzee attended the 8th symposium of the European Association of Acarologists (EURAAC) in Valencia, Spain in July. There were 140 delegates from 33 countries. Topics included mite taxonomy, ecology, biological control, climate change, invasions and forensics. The title of Lizel’s presentation was Distribution patterns of microarthropods on sub-Antarctic Marion Island across altitude and aspect: climate change implications.
From left: Drs Lizel Hugo-Coetzee (South Africa), Andrea Lienhard (Austria), Julia Jagersbacher-Baumann (Austria), Satoshi Shimano (Japan) and Tobias Pfingstl (Austria).
Mites in Art
12 December 2014
Oliewenhuis Art Museum Education Officer, Yolanda de Kock, came up with the brilliant idea to “pair” departments at the National Museum with contemporary South African artists to observe the investigations, results and activities of researchers and use this information as inspiration to create artworks.
Oribatid mites found their way to two artworks – one by Ben Botma, “Tussen-in” (scraperboard and digital print), and one by Elrie Joubert, who made digital prints from light microscope images. For this project, soil was collected from the garden of Oliewenhuis Art Museum, and the oribatids were extracted, identified and photographed.
The project of combining research, education and exhibition fits the mission of the National Museum perfectly: to provide heritage resources and an enjoyable experience to all people through quality research, conservation, education and exhibitions.
The Inquisitive Mind: Science and Imagination can be viewed at Oliewenhuis Art Museum until 1 February 2015.
Second workshop of the Soil Ecosystem Research Group
19 September 2013
Louise Coetzee and Lizel Hugo-Coetzee of the Department of Acarology attended the second meeting of the Soil Ecosystem Research Group (SERG), established in 2011, aiming to form a network to improve communication and develop links between researchers involved in studies of soil biota. The second workshop of SERG was held during the 18th Congress of the Entomological Society of Southern Africa (ESSA XVIII) from 1 to 3 July at Potchefstroom. Progress on a paper to review the status of knowlegde regarding soil biota in South Africa was discussed. The paper is collaboratively drafted with researchers from various institutions and universities, all working on taxonomy of soil biota. Integrative sampling techniques and funding opportunities were also discussed.
EURAAC 2012
21 August 2012
Louise Coetzee attended the 7th symposium of the European Association of Acarologists (EURAAC), held from 9 – 13 July 2012 at the University of Natural Resources and Life Sciences (BOKU), Vienna, Austria. Louise presented a paper “An overview of the genus Afroleius (Acari, Oribatida)” in which certain character states not previously noticed in this genus were illustrated and a reinterpretation of other characters were discussed.
Apart from the scientific content of a conference, it is also a valuable experience to make social contact with colleagues from all over the world.
Louise Coetzee and Prof. Reinhardt Schuster, retired acarologist from Graz, Austria.
Soil Ecology Workshop
21 July 2011
A soil ecology workshop was held on 6 July 2011 during the 17th Congress of the Entomological Society of Southern Africa, held at the University of the Free State from 3 – 6 July. The workshop was attended by taxonomists and other researchers concerned with soil biota. Presentations were made by a number of specialists working on various ecologically important soil dwellers. The message from this workshop was clear: the shortage of taxonomists in these groups is severe. A soil ecology research group was established in order to find measures to address this problem.
Louise Coetzee presented a talk “Mites in Soil” pointing out the ecological role of oribatid (soil) mites as well as the taxonomic status of Oribatida in South Africa. Dr Lizel Hugo-Coetzee presented a poster (co-authored by Dr Nico Avenant) “Soil mite (Acari: Oribatida) response to prescribed fire in a central South African grassland”.
Attendants at the inaugural meeting of the Soil Ecology Research Group (Louise Coetzee 2nd row, second from left; Lizel Hugo-Coetzee 2nd row, third from left).
International Congress of Acarology
6 October 2010
Dr Lizel Hugo-Coetzee from the Acarology Department, National Museum, Bloemfontein, attended the 13th International Congress of Acarology from 23-27 August, 2010, in Recife, Brazil, where she reported on the effect of fire on soil mite communities in Free State grasslands.
The Congress was attended by 380 delegates from 41 countries. Other South Africans present at the Congress were Dr Eddie Ueckermann (Plant Protection Research Institute, Agricultural Research Council), Prof. Pieter Theron (School of Environmental Sciences & Development, North West University), Dr Louwrens Tiedt (Laboratory of Electron Microscopy, North West University) and Dr Jacob den Heyer (Department of Zoology & Entomology, University of the Free State.
Dr Lizel Hugo-Coetzee (4th from left) with participants at the Acarology Congress, Recife, Brazil
Principal Museum Scientist and Head of Department
Lizel Hugo-Coetzee PhD lhugo@nasmus.co.za
Lizel Hugo-Coetzee obtained her BSc, BSc Hons and MSc at the University of the Free State, majoring in Entomology. She obtained her PhD in Conservation Ecology at Stellenbosch University in 2006. Her thesis title was: Spatial patterns in the microarthropod community associated with Azorella selago (Apiaceae) on the sub-Antarctic Prince Edward Islands. Lizel spent a year on Marion Island collecting data for this degree. She started working at the National Museum in January 2006 as a Museum Scientist. Her main interest is the taxonomy of oribatid mites in South Africa, as well as some ecological aspects of oribatid mite communities. Lizel is a Research Associate of the Department of Zoology and Entomology at the University of the Free State. She is also a section editor for the journal, Systematic and Applied Acarology. Lizel has been appointed Head of Department in January 2017.
The objective of this Department is the taxonomic research of Oribatida (Acari) in order to apply our knowledge to control and sustain the condition of soil and the environment. Our main function is to describe the unknown oribatids of South Africa.
Current projects
Lizel Hugo-Coetzee
New species of the Family Oppiidae in South Africa.
Please contact the Acarology department for papers not available from this website.
E.A. Hugo-Coetzee lhugo@nasmus.co.za
Hugo-Coetzee, E.A.2017. New Oppiidae (Acari: Oribatida) from Vernon Crookes Nature Reserve, South Africa. Zootaxa 4311(2): 211-232.
Hugo-Coetzee, E.A.; Lotfollahi, P.; Abbasi-Kalo, A.; Movahedzade, E.2017. A new species of Oxyoppia (Oxyoppiella) (Acari, Oribatida, Oppiidae) from Iran. Turkish Journal of Zoology 41: 630-637.
Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017. Oribatid mites (Acari, Oribatida) inhabiting nests of the termite Trinervitermes trinervoides (Sjöstedt) in the Franklin Game Reserve (Bloemfontein, South Africa), with description of a new species of the genus Ceratobates (Tegoribatidae). Systematic and Applied Acarology 22(10): 1715-1732.
Ermilov, S.G.; Hugo-Coetzee, E.A.;Theron, P.D.2017. New and interesting moss mites (Acari, Oribatida) from South Africa, with description of two new species. Systematic and Applied Acarology22(10): 1560-1573.
Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017. A new genus of the mite family Scutacaridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: Termitidae) from South Africa. Zootaxa 4258(5): 462-476.
Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Contribution to the knowledge of oribatid mites of the family Lohmanniidae (Acari: Oribatida) from South Africa. Systematic and Applied Acarology 22(5): 666-682.
Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017.Two new species of the genus Ausoribula(Acari, Oribatida, Oribatulidae) from termitaries of South Africa. Acarologia 57(3): 643-650.
Khaustov, A.A.; Hugo-Coetzee, E.A.; Ermilov, S.G. 2017.A new genus and two new species of Pygmephoridae (Acari: Heterostigmata) associated with Trinervitermes trinervoides (Isoptera: termitidae) from South Africa. Systematic and Applied Acarology 22(4): 484-493.
Ermilov, S.G.; Hugo-Coetzee, E.A.; Khaustov, A.A. 2017. Coetzeella navalensis gen. nov. sp. nov. (Acari; Oribatida; Oppiidae) from South Africa.Systematic & Applied Acarology 22(3): 403-409.
Ahaniazad, M., Bagheri, M., Akrami, M.A. & Hugo-Coetzee, E.A. 2017. Ameroid mites (Acari; Oribatida) from northwest of Iran with description of a new species of Ctenobelba Balogh, 1943. Systematic & Applied Acarology 22(1): 74–84.
Hugo-Coetzee, E.A. 2016. New species of Ramusella (Oribatida, Oppiidae) from South Africa with additional notes on R. filamentosa (Mahunka) and R. filigera(Mahunka) Systematic & Applied Acarology,21(2): 227–244
Janion-Scheepers, C. et al. 2016. Soil biota in a megadiverse country: Current knowledge and future research directions in South Africa. Pedobiologia 59: 129-174.
Coetzee, L. 2015. Key to the species of Afroleius Mahunka, 1984 (Acari, Oribatida, Punctoribatidae), recombination of A. polygonatus (Mahunka, 1985), description of A. lucidus sp. nov. and discussion of A. undulatus (Balogh, 1959). Systematic & Applied Acarology 20(4): 383–398. view link
Hugo-Coetzee, E.A. 2014. A new species of Adrodamaeus (Acari, Oribatida, Gymnodamaeidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 30(6): 87-99. PDF 1.5 MEG
Coetzee, L. 2014. Rare new species of the genus Afroleius Mahunka, 1984 (Acari, Oribatida, Mycobatidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 30(5): 71-85. PDF 1.8 MEG
Coetzee, L. 2014. Afroleius floridus (Mahunka, 1985) comb. nov. and three new Afroleius Mahunka, 1984 species (Acari: Oribatida: Mycobatidae) from South Africa. Zootaxa (4): 553-573.
Hugo-Coetzee, E.A. 2014. New Oppiidae (Acari: Oribatida) from the Golden Gate Highlands National Park in South Africa. Zootaxa 3884(6): 533-552.
Coetzee, L. 2013. New species of the genus Afroleius Mahunka, 1984 (Acari, Oribatida, Mycobatidae) from South Africa. Acta Zoologica Academiae Scientiarum Hungaricae 59(4): 307-319. PDF 1.9 MEG
Coetzee, L. & Tiedt, L.R. 2013. Overview of the genus Afroleius Mahunka, 1984 Acari, Oribatida). In: Schausberger, P. (ed.) Acari in a changing world: Proceedings of the 7th Symposium of EURAAC, Vienna, 2012. Acarologia 53(2):163-173. PDF 1.1MEG
Hugo-Coetzee, E.A. 2013. New species of Aleurodamaeus Grandjean, 1954 (Oribatida: Aleurodamaeidae) from South Africa. Zootaxa 3670(4): 531-556.
Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species of Nothrus (Acari: Oribatida: Nothridae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 28(2): 25-40. PDF 850 KB
Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Valbehanella freestatensis gen. nov., sp. nov. (Acari, Oribatida) from South Africa. Systematic & Applied Acarology 17(4): 407-416.
Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Oribatid mites of the genus Eremulus Berlese, 1908 (Acari: Oribatida: Eremulidae) from South Africa. African Invertebrates 53(2): 559-569. PDF 860KB
Schuster, R.K. & Coetzee, L. 2012. Cysticercoids of Anoplocephala magna (Eucestoda: Anoplocephalidae) experimentally grown in oribatid mites (Acari: Oribatida). Veterinary Parasitology. view link
Coetzee, L. & Weigmann, G. 2012. Systematic revision of the genus Maudheimia Dalenius, 1958 (Acari: Oribatida). Zootaxa 3295: 65-68.
Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. The oribatid mite genus Nothrus Koch, 1836 (Acari: Oribatida: Nothridae) of South Africa, including a key to African species. Zootaxa 3243: 29-51.
Ermilov, S.G. & Hugo-Coetzee, E.A. 2012. Two new species from South Africa, with remarks on generic diagnosis of Licnodamaeolus Covarrubias, 1998 and taxonomic status of Nacunansella Fernandez & Cleva, 1998 (Acari: Oribatida: Licnodamaeidae). Zootaxa 3167: 32-44.
Hugo-Coetzee, E.A. & Avenant, N.L. 2011. The effect of fire on soil oribatid mites (Acari: Oribatida) in a South African grassland. Zoosymposia 6: 210-220.
Ermilov, S.G., Hugo-Coetzee, E.A. & Khaustov, A.A. 2011. Morphology of juvenile instars of Neoliodes terrestris (Wallwork, 1963) and N. ionicus Sellnick, 1931 (Acari: Oribatida: Neoliodidae). Annales Zoologici (Warszawa) 61(4): 817-830.
Hugo-Coetzee, E.A. 2011. Three new species of Austrocarabodes (Oribatida: Carabodidae) and notes on Austrocarabodes pinnatus Mahunka, 1986, from South Africa. Zootaxa 3011: 1-15.
Hugo-Coetzee, E.A. 2010. Two new species of Austrocarabodes (Acari: Oribatida: Carabodidae) from South Africa: A. mahunkai and A. lineasetosa. Navorsinge van die Nasionale Museum, Bloemfontein26(2): 45-58. PDF 1 MB
Hugo, E.A. 2010. Two new species of Gymnodamaeidae (Acari: Oribatida) from South Africa. International Journal of Acarology 36(3): 199-210.
Coetzee, L. 2010. Species or morphological variation? A multivariate morphometric analysis of Afroleius simplex (Acari, Oribatida, Haplozetidae). M.W. Sabelis & J. Bruin (eds). Trends in Acarology. pp. 267-269. PDF 926KB
Sinclair, W., Coetzee, L. & Joubert, G. 2010. House-dust mite species in Bloemfontein, South Africa. South African Medical Journal 95(3): 630-634. PDF 340KB
Hugo, E.A. 2008. Three new species of Austrocarabodes (Oribatida: Carabodidae) from South Africa. Zootaxa 1844: 25-36.
Coetzee, L. 2007. The genus Afroleius Mahunka (Acari, Oribatida, Haplozetidae) 1. Redescriptions of A. deformis, A. minor and A. simplex. Navorsinge van die Nasionale Museum, Bloemfontein 23(3): 101-118. PDF 1,6MEG.
Akrami, M.A. & Coetzee, L. 2007. Mabulatrichus iranicus (Acari: Oribatida: Zetomotrichidae): a new species from Iran. Systematic and Applied Acarology 12: 245-252. PDF 93KB
Hugo, E.A., Chown, S.L. & McGeoch, M.A. 2006. The microarthropods of sub-Antarctic Prince Edward Island: a quantitative assessment. Polar Biology, 30(1):109-119 PDF 208KB
Grobler, L., Bayram, S. & Cobanoglu, S. 2005. Two new records of Oribatula (Zygoribatula) species (Acari: Oribatida) from Turkey, with redescriptions. Zoological Science 22:1347-1351.
Hugo, E.A., McGeoch, M.A., Marshall, D.J. & Chown, S.L. 2004. Fine scale variation in microarthropod communities inhabiting the keystone species Azorella selago on Marion Island. Polar Biology, 27(8):466-473. PDF 201KB
Yamamoto, Y. & Coetzee, L. 2004. The Trimalaconothrus species from South Africa (Acari: Oribatida: Malaconothridae). Acta Arachnologica 53(1): 13-25. PDF 3.6 MEG
Grobler, L., Bayram, S., & Cobanoglu, S. 2004. Two new species and new records of Oribatid mites from Turkey. International Journal of Acarology 30(4): 351-358.
Yamamoto, Y. & Coetzee, L. 2003. The Malaconothrus species from South Africa and Lesotho (Acari: Oribatida: Malaconothridae). Acta Arachnologica 52(2): 113-126. PDF 3.4 MEG
Grobler, L., Ozman, S.K. & Cobanoglu, S. 2003. The genera Liacarus, Stenoxenillus and Xenillus (Oribatida: Gustavioidea) from Turkey. Acarologia 43(1): 133-149.
Grobler, L. 2003. The genus Furcoppia (Acari, Oribatida) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 19(7): 161-179
Coetzee, L. & Brink, J.S. 2003. Fossil oribatid mites (Acari, Oribatida) from the Florisbad Quaternary deposits, South Africa. Quaternary Research 59: 246-254. PDF 3,3MEG
Coetzee, L. & Marshall, D.J., 2003. A new Halozetes species (Acari: Oribatida: Ameronothridae) from the marine littoral of southern Africa. African Zoology 38(2): 327-331. PDF 427KB
Coetzee, L. 2003. A new genus and species Floritrichus louisbothai (Acari, Oribatida, Oripodoidea, Zetomotrichidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 9(5):93-100. PDF 335KB
Coetzee, L. 2002. Preliminary report on fossil oribatid mites from the Florisbad quaternary research station, South Africa. F. Bernini et al. (eds). Acarid Phylogeny an Evolution Adaptations in mites and ticks, pp. 41-44. PDF 1,3 MEG
Coetzee, L. 2001. Lohmanniidae species (Acari, Oribatida) from the Holocene Deposits at Florisbad, South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 17(5):125-134. PDF 237KB
Coetzee, L. 2001. New species of the Family Lohmanniidae (Acari, Oribatida) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 17(3):53-67. PDF 327KB
Heyne, H., Ueckermann, E.A. & Coetzee. L. 2001. First report of a parasitic mite, Leptotrombidium subquadratum (Lawrence) (Acari: Trombiculidae, Trombiculinae), from dogs and children in the Bloemfontein area, South Africa. Journal of the South African Veterinary Association. 72(2):105-106. PDF 1,3 MEG
Coetzee, L. 2000. A new subspecies of Halozetes belgicae (Acari, Oribatida, Ameronothridae) from the Marine supra-littoral of the sub-Antarctic Marion Island. Acarologia. 15(4):439-447. PDF 2MEG
Grobler, L & Skubala, P. 2000. Description of Oribatula dentata, sp. nov. (Oribatulidae, Oribatida) from Spain and complementary data on O. longolamellata and O. macrostega. Acarologia 15(3): 343-353.
Marshall, D.J. & Coetzee, L. 2000. Historical biogeography and ecology of a Continental Antarctic mite genus, Maudheimia (Acari, Oribatida): evidence for a Gondwanan origin and Pliocene-Pleistocene speciation. Zoological Journal of the Linnean Society 129:111-128. PDF 2,6 MEG
Bayartogtokh, B., Grobler, L. & Cobanoglu, S. 2000. A new species of Punctoribates (Acari: Oribatida: Mycobatidae) collected from mushrooms in Turkey, with remarks on the taxonomy of the genus. Navorsinge van die Nasionale Museum, Bloemfontein 16(2):18-32.
Schuster, R., Coetzee. L. & Putterill, J.F. 2000. Oribatid mites (Acari, Oribatida) as intermediate hosts of tapeworms of the Family Anoplocephalidae (Cestoda) and the transmission of Moniezia expansa cysticercoids in South Africa. Onderstepoort Journal of Veterinary Research 67:49-55. PDF 1.4 MEG
Grobler, L. 2000. Megatrichobates Gen. Nov. (Acari, Oribatida, Oribatulidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 16(1):1-14.
Marshall, D.J., Gremmen, N.J.M., Coetzee, L., O'Connor, B.M., Pugh, P.J.A., Theron, P.D. & Ueckermann, E.A. 1999. New records of Acari from the sub-Antarctic Prince Edward Islands. Polar Biology 21:84-89. PDF 2,0 MEG
Grobler, L. 1998. Subfamily Pseudotocepheinae (Acari, Oribatida, Otocepheidae): Constrictocepheusgen. nov. and new Pseudotocepheus species from South Africa Navorsinge van die Nasionale Museum, Bloemfontein 14(1): 1-44.
Coetzee, L. 1997. The Antarctic mite genus Maudheimia (Acari, Oribatida). Navorsinge van die Nasionale Museum, Bloemfontein 13 (3): 101-135. PDF 4,2 MEG
Grobler, L. 1997. The Subfamily Pseudotocepheinae subfam. nov. (Acari, Oribatida, Otocepheidae). Navorsinge van die Nasionale Museum, Bloemfontein 13(1): 1-44.
Grobler, L. 1995. Two Otocepheid genera (Acari, Oribatida) from South Africa: Longocepheus Balogh & Mahunka and Trichocepheus Balogh & Mahunka. Navorsinge van die Nasionale Museum, Bloemfontein 11(10): 261-300.
Grobler, L. 1995. New species of the genus Plenotocepheus Hammer, 1966 (Acari, Oribatida, Otocepheidae) from South Africa II. Navorsinge van die Nasionale Museum, Bloemfontein 11(5): 102-118.
Grobler, L. 1995. New species of the genus Plenotocepheus Hammer, 1966 (Acari, Oribatida, Otocepheidae) from South Africa I. Navorsinge van die Nasionale Museum, Bloemfontein 11(2): 13-36.
Coetzee, L. 1995. A new species of the genus Saltatrichus Coetzee, 1993 (Acari, Oribatida, Zetomotrichidae) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 11(1): 1-12. PDF 1,1 MEG
Grobler, L. 1994. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa III. Zygoribatula contracta spec. nov. and Zygoribatula setosa Evans, 1953. Navorsinge van die Nasionale Museum, Bloemfontein 10(6): 261-279.
De Wet, L. 1993. New species of the genus Hermannia Nicolet, 1855 from South Africa II. (Acari, Oribatida, Hermanniidae). Navorsinge van die Nasionale Museum, Bloemfontein 9(9): 266-287.
Grobler, L. & Kok, D.J. 1993. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa II. New and existing species. Navorsinge van die Nasionale Museum, Bloemfontein 9(6): 182-212.
Coetzee, L. 1993. New genera and species of the family Zetomotrichidae Grandjean, 1954 (Acari, Oribatida, Oripodoidea) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 9(5): 133-179. PDF 4,1 MEG
Grobler, L. 1993. Species of the genus Zygoribatula Berlese, 1916 (Acari, Oribatida, Oribatulidae) from South Africa I. New species. Navorsinge van die Nasionale Museum, Bloemfontein 9(3): 49-76.
De Wet, L. 1993. New species of the genus Hermannia Nicolet, 1855 from South Africa I. (Acari: Oribatida: Hermanniidae). Navorsinge van die Nasionale Museum, Bloemfontein 9(2): 21-48.
Grobler, L. 1991. A new Brasilobates Perez-Inigo & Baggio, 1980 from South Africa (Acari: Oribatei: Oribatuloidea: Xylobatidae). Navorsinge van die Nasionale Museum, Bloemfontein 7(5): 73-95.
Grobler, L. 1989. New South African species of the genus Eupelops Ewing, 1917 (Acari: Oribatei: Phenopelopoidea: Phenopelopidae). Navorsinge van die Nasionale Museum, Bloemfontein 6(5): 151-201.
Coetzee, L. 1989. The genus Zetorchestes Berlese, 1888 (Acari, Oribatei, Zetorchestidae) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 6(6): 203-221. PDF 1,5 MEG
Coetzee, L. 1987. The South African Lamellareidae Balogh, 1972 (Acari: Oribatei). Navorsinge van die Nasionale Museum, Bloemfontein 5(13): 325-353. PDF 2,9 MEG
Engelbrecht, C.M. 1986. New Oribatid taxa and distribution records predominantly from Southern Africa. (Acari: Oribatei: Oribatelloidea). Navorsinge van die Nasionale Museum, Bloemfontein 5(9): 170-250.
Engelbrecht, C.M. 1975. Die Suid-Afrikaanse Eupelopoidea Balogh, 1963. Navorsinge van die Nasionale Museum, Bloemfontein 3(5): 89-108.
Engelbrecht, C.M. 1975. New Ameronothroid (Oribatei, Acari) taxa from the Republic of South Africa and the Islands Gough and Marion. Navorsinge van die Nasionale Museum, Bloemfontein 3(4): 53-88.
Engelbrecht, C.M. 1974. The genus Hydrozetes Berlese, 1902. (Oribatei: Acari) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 3(3): 41-49.
Engelbrecht, C.M. 1974. The genus Passalozetes Grandjean, 1932 (Oribatei: Acari) in South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 3(2): 29-38.
Engelbrecht, C.M. 1974. The genus Halozetes (Oribatei: Acari) on Marion Island. Navorsinge van die Nasionale Museum, Bloemfontein 3(1): 1-25.
Engelbrecht, C.M. 1973. Magoebazetes, a new microzetid genus (Microzetidae: Oribatei) from South Africa. Acarologia 14(4): 681-695.
Engelbrecht, C.M. 1973. South African Galumnoidea (Oribatei: Acari): New taxa and records. Navorsinge van die Nasionale Museum, Bloemfontein 2(11): 394-415.
Engelbrecht, C.M. 1972. A new South African microzetid genus, Afrozetes (Microzetidae: Oribatei). Acarologia 14(3): 484-496.
Engelbrecht, C.M. 1972. Ctenogalumna moresonensis sp. n. and Pilogalumna bloemfonteinensis sp. n., two new South African species of the subfamily Allogalumninae Balogh, 1960 (Galumnidae: Oribatei). Acarologia 14(3): 497-510.
Engelbrecht, C.M. 1972. South African species of the genus Trichogalumna Balogh, 1961 (Galumnidae: Oribatei). Acarologia 14(2): 268-280.
Engelbrecht, C.M. 1972. Galumnids from South Africa. Acarologia 14(1): 109-140.
Engelbrecht, C.M. 1972. Dimidiogalumna villiersensis (Galumnidae: Oribatei), a new South African galumnid genus and its type species. Acarologia 14(1): 141-149.
Engelbrecht, C.M. 1972. A new microzetid genus and species Schalleriella grabouwensis(Microzetidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 282-285.
Engelbrecht, C.M. 1972. Microzetes auxiliaris orkneyensis s. sp. n. and Microzetes natalensis sp. n. (Microzetidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 270-281.
Engelbrecht, C.M. 1972. Two new species of the genus Rhopalozetes Balogh, 1961 Rhopalozetes georgensis sp. n. and Rhopalozetes witfonteinensis sp. n. (Microzetidae: Oribatei). Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 263-269.
Engelbrecht, C.M. 1972. Two new species of the genus Galumna Von Heyden, 1826 (Galumnidae: Oribatei) from South Africa. Navorsinge van die Nasionale Museum, Bloemfontein 2(9): 244-262.
Engelbrecht, C.M. 1969. Some South African species of the genus Galumna Von Heyden, 1826 (Acari: Galumnidae). Journal of the Entomological Society of Southern Africa 32(1): 99-122.
Articles from Culna 60 onwards available at culna-downloads/
Hugo-Coetzee, L. 2016. Mites at the beach. Culna 71: 14-15
Coetzee, L. 2015. Die alomteenwoordige myte. Culna 70: 20-21.
Hugo-Coetzee, L. 2014. Soil mites on guard. Culna 69:24-25.
Coetzee, L. 2013. Grondmyte en die stryd teen waterhiasinte. Culna 68:11-13.
Hugo-Coetzee, L. 2013. What's in a name? From Hitler to the vuvuzela. Culna 68: 20-21.
Hugo-Coetzee, L. 2012. Die kortafstand ... vir insekte. Culna 67: 26-27
Coetzee, L. 2011. Myte - het ons hulle nodig? Culna 66: 12-14
Hugo-Coetzee, L. 2011. Living Soil. Culna 66: 22-23
Coetzee, L. 2009. Ongewone bedmaats - huisstofmyte. Culna 64: 29-39
Hugo, L. 2009. Describing species - how it is done. Culna 64: 19-20
Hugo, L. 2008. Goggas in die water. Culna 63: 17-18
Hugo, L. 2007. The sex life of soil mites. Culna 62: 16-17
Coetzee, L. 2006. The Secretarybird - majestic hunter of the plains. Culna 61: 30-31
Hugo, L. 2006. Myte - die agtbenige "ysbere" van Antarktika. Culna 61: 16-17
Coetzee, L. 2005. Chiggers - are your dogs biting their paws? Culna 60: 19-20
Coetzee, L. 2004. What makes an owl? Culna 59: 17-19
Coetzee, L. 2000. Oribatid mites found in the Quaternary sediments at Florisbad. Culna 55: 21
Coetzee, L. 1998. Wat gaan binne in die grond aan? Culna 53: 16
Coetzee, L. 1984. Oribatei. National Museum News 27: 26-27
Taxonomic research is focussed mainly on the Oribatida.
The collection currently consists of 27 220 oribatid samples (each containing a varying number of specimens), collected from 4 580 localities. The samples were collected from a wide range of habitats in South Africa, Gough Island and the sub Antarctic Islands Marion and Prince Edward. The specimens are stored in 70% Alcohol + 5% Glycerol.
The holotype collection consists of 223 Oribatida, 19 Astigmata (Analgoidea), collected from birds in South Africa, and 13 Prostigmata (Hygrobatoidea).
The collection of Oribatida holotypes which belongs to the Ditsong National Museum of Natural History, Pretoria, (formerly the Transvaal Museum), are now at the National Museum, Bloemfontein, on a long term loan. This material comprises oribatids collected from South Africa and studied by Dr. Sandor Mahunka.
Astigmata Holotypes
Analgidae | Anagles vanasi Mironov & Kopij, 1996 |
Proctophyllodidae | Montesauria belozerovi Mironov & Kopij, 1996 |
Montesauria emberizae Mironov & Kopij, 1996 | |
Montesauria jesionowskii Mironov & Kopij, 1997 | |
Montesauria punctata Mironov & Kopij, 1996 | |
Montesauria tetralobula Mironov & Kopij, 1996 | |
Pedanodectes marginatus Mironov & Kopij, 1997 | |
Proctophyllodes caffer Mironov & Kopij, 1996 | |
Proctophyllodes leptodiscus Mironov & Kopij, 1996 |
Pteronyssidae | Metapteronyssus angolensis Mironov & Kopij, 2000 |
Pteroherpus africanus Mironov & Kopij, 2000 | |
Pteronyssoides nectariniae Mironov, 2001 | |
Pteronyssoides promeropis Mironov & Kopij, 2000 | |
Sturnotrogus creatophorae Mironov & Kopij, 2000 |
Trouessartiidae | Trouessartia angolensis Mironov & Kopij, 2000 |
Trouessartia creatophorae Mironov & Kopij, 1996 | |
Trouessartia daberti Mironov & Kopij, 2000 | |
Trouessartia priniae Mironov & Kopij, 2000 | |
Trouessartia pytiliae Mironov & Kopij, 2000 |
Oribatida Holotypes
Achipteriidae | Novoribatella minutisetarum Engelbrecht, 1986 |
Novoribatella pentasetarum Engelbrecht, 1986 |
Aleurodamaeidae | Aleurodamaeus salvadordalii Hugo-Coetzee, 2013 |
Aleurodamaeus vicinus Hugo-Coetzee, 2013 | |
Aleurodamaeus angelae Hugo-Coetzee, 2013 | |
Aleurodamaeus niedbalai Hugo-Coetzee, 2013 | |
Aleurodamaeus minutus Hugo-Coetzee, 2013 | |
Aleurodamaeus woasi Hugo-Coetzee, 2013 | |
Aleurodamaeus prominens Hugo-Coetzee, 2013 |
Ameronothridae | Alaskozetes bouvetoyaensis Van Pletzen & Kok |
Halozetes belgicae mickii Coetzee, 2000 | |
Halozetes edwardensis Van Pletzen & Kok | |
Halozetes fulvus Engelbrecht, 1975 | |
Halozetes marinus devilliersi Engelbrecht, 1974 | |
Halozetes marionensis Engelbrecht, 1974 |
Aquanothridae | Aquanothrus montanus Engelbrecht, 1975 |
Carabodidae | Austrocarabodes hendriksi Hugo, 2008 |
Austrocarabodes luciensis Hugo, 2008 | |
Austrocarabodes vernoncrookensis Hugo, 2008 | |
Austrocarabodes mahunkai Hugo-Coetzee, 2010 | |
Austrocarabodes lineasetosa Hugo-Coetzee, 2010 | |
Austrocarabodes nortoni Hugo-Coetzee, 2011 | |
Austrocarabodes longisetosus Hugo-Coetzee, 2011 | |
Austrocarabodes crassimarginatus Hugo-Coetzee, 2011 |
Crotonidae | Crotonia dicella Colloff, 1990 |
Crotonia ecphyla Colloff, 1990 | |
Crotonia lyrata Colloff, 1990 | |
Crotonia pauropelor Colloff, 1990 |
Cymbaeremaeidae | Scapheremaeus pundamiliaensis Engelbrecht, 1975 |
Scapheremaeus rustenburgenesis Engelbrecht, 1975 |
Eremaeozetidae | Eremaeozetes capensis Schatz, 2001 |
Eremaeozetes louisae Schatz, 2003 | |
Eremaeozetes nasutus Schatz, 2003 | |
Mahunkaia tricornis Schatz, 2002 |
Eremulidae | Eremulus spindleformis Ermilov & Hugo-Coetzee, 2012 |
Eremulus southafricanensis Ermilov & Hugo-Coetzee, 2012 |
Euphthiracaridae | Acrotritia minuta Niedbała, 2006 |
Euphthiracarus rectus Niedbała, 2006 | |
Euphthiracarus scuticus Niedbała, 2006 |
Galumnellidae | Galumnella pilosa Engelbrecht, 1972 |
Galumnidae | Ctenogalumna moresonensis Engelbrecht, 1972 |
Dimidiogalumna villiersensis Engelbrecht, 1972 | |
Galumna capensis capensis Engelbrecht, 1969 | |
Galumna capensis dissimilis Engelbrecht, 1969 | |
Galumna comparabilis Engelbrecht, 1969 | |
Galumna dimidiata Engelbrecht, 1969 | |
Galumna nuda Engelbrecht, 1969 | |
Galumna rostrata Engelbrecht, 1969 | |
Galumna tricuspidata Engelbrecht, 1969 | |
Pergalumna elongata Engelbrecht, 1972 | |
Pergalumna incomperta Engelbrecht, 1972 | |
Pergalumna magnipora capensis Engelbrecht, 1972 | |
Pilogalumna bloemfonteinensis Engelbrecht, 1972 | |
Pilogalumna kimberleyensis Engelbrecht, 1972 | |
Pilogalumna variabilis Engelbrecht, 1972 | |
Sacculogalumna saccularis Engelbrecht1973 | |
Trichogalumna pumularis Engelbrecht, 1972 | |
Trichogalumna punctata Engelbrecht, 1972 |
Gymnodamaeidae | Arthrodamaeus johanni Hugo, 2009 |
Gymnodamaeus deswardti Hugo, 2009 | |
Adrodamaeus ermilovi Hugo-Coetzee, 2014 |
Hermanniidae | Hermannia comparabilis De Wet, 1993 |
Hermannia engelbrechti De Wet, 1993 | |
Hermannia exobothridialis De Wet, 1993 | |
Hermannia louisiae De Wet, 1993 | |
Hermannia natalensis De Wet, 1993 | |
Hermannia nathanaeli De Wet, 1993 | |
Hermannia notogastralis De Wet, 1993 | |
Hermannia phylliformis De Wet, 1993 | |
Hermannia setiformis De Wet, 1993 | |
Hermannia transvaalensis De Wet, 993 |
Hydrozetidae | Hydrozetes capensis Engelbrecht, 1975 |
Lamellareidae | Lamellarea ardua Kok, 1968 |
Lamellarea digitata Kok, 1968 | |
Lamellarea forceps Kok, 1968 | |
Lamellarea quadrata Coetzee, 1986 | |
Microlamellarea engelbrechti Coetzee, 1986 |
Licnodamaeidae | Licnodamaeolus asetosus Ermilov & Hugo-Coetzee, 2012 |
Licnodamaeolus erfenisdamensis Ermilov & Hugo-Coetzee, 2012 |
Lohmanniidae | Annectacarus eksteeni Coetzee, 2001 |
Papillacarus brinki Coetzee, 2001 | |
Paulianacarus barlowi Coetzee, 2001 | |
Paulianacarus grobleri Coetzee, 2001 |
Malaconothridae | Malaconothrus longidorsus Yamamoto & Coetzee, 2004 |
Malaconothrus minimus Yamamoto & Coetzee, 2004 | |
Malaconothrus stigmatus Yamamoto & Coetzee, 2004 | |
Trimalaconothrus binodulus Yamamoto & Coetzee, 2004 | |
Trimalaconothrus duaculeus Yamamoto & Coetzee, 2004 | |
Trimalaconothrus obesus Yamamoto & Coetzee, 2004 | |
Trimalaconothrus punctum Yamamoto & Coetzee, 2004 | |
Trimalaconothrus rectus Yamamoto & Coetzee, 2004 |
Maudheimiidae | Maudheimia marshalli Coetzee, 1997 |
Maudheimia tanngardenensis Coetzee, 1997 |
Mesoplophoridae | Mesoplophora iuvenalis Niedbała, 2006 |
Mesoplophora pertenuis Niedbała, 2006 | |
Mesoplophora setulosa Niedbała, 2006 |
Metrioppiidae | Furcoppia dentata Grobler, 2003 |
Furcoppia longiseta Grobler, 2003 | |
Furcoppia minuta Grobler, 2003 |
Microzetidae | Afrozetes champaignensis Engelbrecht, 1972 |
Afrozetes magoebaensis capensis Engelbrecht, 1972 | |
Afrozetes magoebaensis magoebaensis Engelbrecht, 1972 | |
Afrozetes natalensis Engelbrecht, 1972 | |
Magoebazetes magoebaensis Engelbrecht, 1973 | |
Magoebazetes natalensis Engelbrecht, 1973 | |
Magoebazetes pentasetarum Engelbrecht, 1973 | |
Microzetes auxiliaris orkneyensis Engelbrecht, 1972 | |
Microzetes natalensis Engelbrecht, 1972 | |
Rhopalozetes georgensis Engelbrecht, 1972 | |
Rhopalozetes witfonteinensis Engelbrecht, 1972 | |
Schalleriella grabouwensis Engelbrecht, 1972 |
Nothridae | Nothrus bilongisetosus Ermilov & Hugo-Coetzee, 2012 |
Nothrus monolongisetosus Ermilov & Hugo-Coetzee, 2012 | |
Nothrus louiseae Ermilov & Hugo-Coetzee, 2012 | |
Nothrus engelbrechti Ermilov & Hugo-Coetzee, 2012 |
Oppiidae | Amerioppia africana Kok, 1967 |
Brachioppia longisetosa Kok, 1967 | |
Brachioppia moresonensis Kok, 1967 | |
Brachioppia orkneyensis Kok, 1967 | |
Brachioppia pectinata Kok, 1967 | |
Brachioppia quathlambae Kok, 1967 | |
Oppia falxa Kok, 1967 | |
Oppia parva Kok, 1967 | |
Oppia pletzenae Kok, 1967 | |
Oppia quattuor Kok, 1967 | |
Oppia yodai africana Kok, 1967 | |
Brachioppia louwi Hugo-Coetzee, 2014 | |
Brachioppiella (B) dawidi Hugo-Coetzee, 2014 | |
Brachioppiella (B) goblina Hugo-Coetzee, 2014 | |
Kokoppia mandelai Hugo-Coetzee, 2014 | |
Lanceoppia scytheae Hugo-Coetzee, 2014 | |
Oppiella (Quattoppiella) goldengatensis Hugo-Coetzee, 2014 |
Oribatulidae | Megatrichobates longisetosus Grobler, 2000 |
Megatrichobates striatus Grobler, 2000 | |
Zygoribatula connexa substriata Grobler, 1993 | |
Zygoribatula gracilata Grobler, 1993 | |
Zygoribatula lineaporosa Grobler, 1993 | |
Zygoribatula pennata Grobler, 1993 | |
Zygoribatula robusta Grobler, 1993 | |
Zygoribatula subantarctica Van Pletzen & Kok |
Oribotritiidae | Austrotritia engelbrechti Niedbała, 2006 |
Indotritia cypha Niedbała, 2006 | |
Indotritia didyma Niedbała, 2006 | |
Indotritia eksteeni Niedbała, 2006 | |
Indotritia fusa Niedbała, 2006 | |
Indotritia partita Niedbała, 2006 | |
Indotritia phymatha Niedbała, 2006 | |
Oribotritia deminuta Niedbała, 2006 |
Otocepheidae | Longocepheus globosus Grobler, 1995 |
Plenotocepheus crinitus Grobler, 1995 | |
Plenotocepheus dentatus Grobler, 1995 | |
Plenotocepheus discrepans Grobler, 1995 | |
Plenotocepheus verrucosus Grobler, 1995 | |
Pseudotocepheus aokii Grobler, 997 | |
Pseudotocepheus contractus Grobler, 1997 | |
Pseudotocepheus dentatus Grobler, 1998 | |
Pseudotocepheus lanceolatus Grobler, 1998 | |
Pseudotocepheus promissus Grobler, 1998 | |
Pseudotocepheus shauni Grobler, 1997 | |
Trichocepheus auriculus Grobler, 1995 |
Parakalummidae | Carinogalumna montana Engelbrecht, 1973 |
Passalozetidae | Passalozetes douglasensis Engelbrecht, 1974 |
Passalozetes wolwekransensis Engelbrecht, 1974 |
Phenopelopidae | Eupelops capensis Grobler, 1989 |
Eupelops curtipilus thabazimbiensis Engelbrecht, 1975 | |
Eupelops distinctus Grobler, 1989 | |
Eupelops engelbrechti Grobler, 1989 | |
Eupelops foveolatus Engelbrecht, 1975 | |
Eupelops margatensis Engelbrecht, 1975 | |
Eupelops minutus Grobler, 1989 |
Phthiracaridae | Phthiracarus anakolos Niedbała, 2006 |
Phthiracarus blythedalensis Niedbała, 2006 | |
Phthiracarus densus Niedbała, 2006 | |
Phthiracarus hillcrestensis Niedbała, 2006 | |
Phthiracarus humilis Niedbała, 2006 | |
Phthiracarus kokae Niedbała, 2006 | |
Phthiracarus leliehoekensis Niedbała, 2006 | |
Phthiracarus pertenuis Niedbała, 2006 |
Punctoribatidae | Afroleius crassus Coetzee, 2013 |
Afroleius decurvatus Coetzee, 2013 | |
Afroleius deformatoides Coetzee, 2013 | |
Afroleius amieae Coetzee, 2014 | |
Afroleius inae Coetzee, 2014 | |
Afroleius valerieae Coetzee, 2014 | |
Afroleius natalensis Coetzee, 2014 | |
Afroleius caudatus Coetzee, 2014 | |
Afroleius lucidus Coetzee, 2015 |
Scheloribatidae | Scheloribates durbanensis Van Pletzen, 1963 |
Scheloribates elsi Van Pletzen, 1965 | |
Scheloribates louwi Van Pletzen, 1965 | |
Scheloribates minutus Van Pletzen, 1965 | |
Scheloribates natalensis Van Pletzen, 1963 | |
Scheloribates parvus Van Pletzen, 1963 | |
Scheloribates robustus Van Pletzen, 1963 | |
Scheloribates rustenburgensis Van Pletzen, 1963 | |
Scheloribates tzitzikamaenesis Van Pletzen, 1965 | |
Scheloribates variabilis Van Pletzen, 1965 |
Scutoverticidae | Scutovertex sculperens Kok, 1968 |
Scutoverticosus arcanus Kok, 1968 |
Steganacaridae | Arphthicarus bulbosus Niedbała, 2006 |
Atropacarus brevisetosus Niedbała, 2006 | |
Austrophthiracarus cordylus Niedbała, 2006 | |
Notophthiracarus agulhasensis Niedbała, 2006 | |
Notophthiracarus capevidalensis Niedbała, 2006 | |
Notophthiracarus coetzeeae Niedbała, 2006 | |
Notophthiracarus cristatus Niedbała, 2006 | |
Notophthiracarus frondeus Niedbała, 2006 | |
Notophthiracarus gongylos Niedbała, 2006 | |
Notophthiracarus grossus Niedbała, 2006 | |
Notophthiracarus knysnaensis Niedbała, 2006 | |
Notophthiracarus korannabergensis Niedbała, 2006 | |
Notophthiracarus mekistos Niedbała, 2006 | |
Notophthiracarus natalensis Niedbała, 2006 | |
Notophthiracarus phyllodes Niedbała, 2006 | |
Notophthiracarus procerus Niedbała, 2006 | |
Notophthiracarus pumilus Niedbała, 2006 | |
Notophthiracarus rhachis Niedbała, 2006 | |
Notophthiracarus serratus Niedbała, 2006 | |
Notophthiracarus tsitsikamaensis Niedbała, 2006 | |
Notophthiracarus vernonensis Niedbała, 2006 | |
Notophthiracarus zululandensis Niedbała, 2006 | |
Protophthiracarus crinatus Niedbała, 2006 | |
Protophthiracarus engelbrechti Niedbała, 2006 |
Tectocepheidae | Tegeocranellus knysnaensis Kok, 1968 |
Tegeocranellus sacchareus Kok, 1968 |
Tegoribatidae | Tegoribates montana Engelbrecht, 1986 |
Tegoribates natalensis Engelbrecht, 1986 | |
Tegoribates nuda Engelbrecht, 1986 |
Unduloribatidae | Valbehanella freestatensis Ermilov & Hugo-Coetzee, 2012 |
Xylobatidae | Brasilobates punctata Grobler, 1991 |
Zetomotrichidae | Demisalto engelbrechti Coetzee, 1993 |
Floritrichus louisbothai Coetzee, 2003 | |
Mabulatrichus dentatus Coetzee, 1993 | |
Saltatrichus magnus Coetzee, 1993 | |
Saltatrichus marshalli Coetzee, 1995 | |
Saltatrichus minus Coetzee, 1993 | |
Saltatrichus robustus Coetzee, 1993 |
Zetorchestidae | Zetorchestes transvaalensis Coetzee, 1989 |
Prostigmata Holotypes
Hygrobatidae | Hygrobates draconicus Smit, 2005 |
Hygrobates inflatipes Smit, 2005 | |
Hygrobates laceratoides Smit, 2005 | |
Hygrobates spathuliferus cederberg Smit, 2005 | |
Atractides capensis Pesic, Smit & Gerecke, 2011 | |
Atractides cooki Pesic, Smit & Gerecke, 2011 | |
Atractides dracomontanus Pesic, Smit & Gerecke, 2011 | |
Atractides meridianus Pesic, Smit & Gerecke, 2011 | |
Atractides neoscutifer Pesic, Smit & Gerecke, 2011 |
Torrenticolidae | Monatractides nigromontanus Goldschmidt & Smit, 2009 |
Monatractides drakenbergensis Goldschmidt & Smit, 2009 | |
Torrenticolidae spatulifra Goldschmidt & Smit, 2009 | |
Torrenticola tsitsikamma Goldschmidt & Smit, 2009 |
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